ALLIANCE CONCEPTSummary: This alliance, found near the Great Lakes and in the southern Appalachian Mountains and northeastern United States, iscomposed of dry-mesic to mesic pine forests. Stands of this alliance are characterized by a moderate to complete tree canopy. Theshrub layer is absent to well-developed, while the herbaceous layer is moderately to poorly developed. Understory vegetation is sparsewhere the canopy is closed, due to the limited amount of light and the duff buildup on the forest floor. The overstory is heavilydominated by coniferous trees, usually Pinus strobus alone but sometimes with Pinus resinosa. Other canopy and subcanopy treesinclude Abies balsamea (in the northern part of this alliance's range), Acer rubrum, Betula papyrifera, Populus tremuloides, and Thujaoccidentalis. The shrub layer typically contains species such as Acer spicatum, Corylus cornuta, Diervilla lonicera, Linnaea borealis,and Vaccinium spp., especially Vaccinium myrtilloides and Vaccinium angustifolium. The herb layer contains species adapted to thedry-mesic nature of stands of this alliance. These include Aralia nudicaulis, Eurybia macrophylla (= Aster macrophyllus), Gaultheriaprocumbens, and Maianthemum canadense.Stands of this alliance are found on loamy sand, sandy loam, loam, and clay loam soils which are typically moderately deep to deep(60-100 cm) except in the Driftless Area where they may be very shallow. The soils are acidic and rarely contain a significant amountof organic material. Stands of this alliance are often found on glacial till or outwash plains, although in northeastern Minnesota theyoccur near lakes and on lower slopes. This alliance can be found on a variety of landscapes, varying from nearly level to rolling acrossmuch of its range to steep slopes in the Driftless Area. In the southern Appalachians these forests occur below 3000 feet (900 m)elevation on upper slopes and ridgetops protected by higher landforms, or as successional forests on abandoned agricultural land.Related Concepts:• Eastern Needleleaf <strong>Forest</strong>s: 95: Great Lakes Pine <strong>Forest</strong> (Pinus) (Kuchler 1964) I• Eastern White Pine: 21 (Eyre 1980) I IA6f. Dry White Pine Ridge <strong>Forest</strong> (Allard 1990) I• Northern Dry-mesic <strong>Forest</strong> (Curtis 1959) I• White Pine BR, RV, CUPL (Pyne 1994) ?• White Pine <strong>Forest</strong> (Schafale and Weakley 1990) ?Classification Comments: Natural Pinus strobus stands occur in mesic gorges of eastern Kentucky over Rhododendron maximum orwith a lush herbaceous stratum. KP 11-99: might these be closer to CEGL007102, in A.127? MP: Pinus strobus occurs as a disjunctspecies in Tennessee's Western Highland Rim (Cheatham and Dickson counties) but occurs in a mixed pine - oak forest community.ALLIANCE DISTRIBUTIONRange: This alliance is found in Michigan, northern Wisconsin, northern and eastern Minnesota, extreme northeastern Iowa, Maine,New Hampshire, North Carolina, South Carolina, Georgia, Tennessee, Kentucky (?), and Virginia. In Canada, it is found in Ontario.Subnations: GA, IA, KY?, ME, MI, MN, NB, NC, NH, NS?, NY, ON, PA, QC?, SC, TN, VA, VT, WI, WVTNC Ecoregions: 46:C, 47:C, 48:C, 49:C, 50:C, 51:C, 59:C, 60:C, 61:C, 62:C, 63:C, 64:CUSFS Ecoregions: 212Aa:CC?, 212Ab:CC?, 212Ba:CCP, 212Bb:CCP, 212Ca:CCC, 212Cb:CCC, 212Da:CCC, 212Ea:CCP,212Eb:CCP, 212Ec:CCC, 212Fa:C??, 212Fb:C??, 212Fc:C??, 212Ga:C??, 212Ha:CCC, 212Hb:CCP, 212He:CCP, 212Hh:CCC,212Hi:CCP, 212Hj:CCC, 212Hl:CCC, 212Hm:CCC, 212Ho:CCC, 212Hp:CCC, 212Hq:CCP, 212Hr:CCC, 212Hs:CCP, 212Ht:CCP,212Hu:CCP, 212Hv:CCP, 212Hw:CCP, 212Hx:CCP, 212Hy:CCP, 212Ia:CCC, 212Ja:CCP, 212Jb:CCC, 212Jc:CCC, 212Je:CCC,212Jf:CC?, 212Jj:CC?, 212Jl:CCC, 212Jm:CCC, 212Jn:CCP, 212Jr:CCC, 212Ka:CCC, 212Kb:CCC, 212La:CCC, 212Lb:CCC,212Lc:CCC, 212Ld:CCP, 212Ma:CCC, 212Mb:CCC, 212Na:CCC, 212Nb:CCC, 212Nc:CCC, 212Nd:CCP, 221Ae:CCC,221Af:CCP, 221Al:CCC, 221Ba:CPP, 221Ha:CCC, 221Hb:CCC, 221Hc:CCC, 221He:CCC, 221Ja:C??, 221Jc:C??, 222En:CCC,222Eo:CCC, 222Ib:C??, 222Ic:C??, 222Id:C??, 222Ie:C??, 222If:C??, 222Lc:CCC, 222Ld:CCC, 222Le:CCC, 222Lf:CCC,222Ma:CCC, 222Mc:CCC, 231:C, M212Ab:CCC, M212Ac:CCC, M212Ae:CCC, M212Ag:CCC, M212Ba:CCC, M212Bc:CCC,M212Cb:CCC, M212Cc:CCC, M212Da:CCP, M212Db:CCP, M212Dc:CCP, M212Ea:CCP, M212Eb:CCP, M212Fa:C??,M212Fb:C??, M221Aa:CCC, M221Ab:CCC, M221Ac:CCC, M221Bb:CCC, M221Bd:CCC, M221Bf:CCP, M221Cb:CPP,M221Cd:CPP, M221Da:CCC, M221Db:CCC, M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Acadia, Blue Ridge Parkway?, Carl Sandburg Home, Great Smoky Mountains, Voyageurs); USFS(Chattahoochee, Cherokee, Daniel Boone, George Washington, Jefferson, Nantahala, Pisgah, <strong>Sumter</strong>)ALLIANCE SOURCESReferences: Allard 1990, Burns and Honkala 1990a, Curtis 1959, DeYoung 1979, DuMond 1970, Eyre 1980, Faber-Langendoen etal. 1996, Govus 1982, Hinkle 1989, Kuchler 1964, MNNHP 1993, Ohmann and Ream 1971, Patterson 1994, Pyne 1994, Schafale andWeakley 1990, Sims et al. 1989, Tobe et al. 1992I.A.8.N.B. PINUS TAEDA FOREST ALLIANCE (A.130)LOBLOLLY PINE FOREST ALLIANCEALLIANCE CONCEPTSummary: This alliance includes both successional forests, following cropping or site conversion, and natural forests in thePiedmont, Cumberlands and Ridge and Valley, and Coastal Plain of the southeastern United States. Other canopy and subcanopyspecies that may be present in successional stands are Liriodendron tulipifera, Acer rubrum, Liquidambar styraciflua, PinusVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>4
virginiana, Juniperus virginiana var. virginiana, Quercus stellata, Quercus velutina, Ulmus rubra, Quercus alba, Nyssa sylvatica,Ulmus alata, Cornus florida, Prunus serotina var. serotina, and Carya spp. Vaccinium spp., especially Vaccinium stamineum, arecommon in these forests. One association in this alliance occurs on barrier islands in the Mid-Atlantic Coastal Plain. Along with thedominant Pinus taeda, canopy associates often include Quercus falcata, Acer rubrum, Prunus serotina var. serotina, and Sassafrasalbidum. The tall-shrub layer is comprised of Morella cerifera (= Myrica cerifera) and Vaccinium formosum. Vines and lianas arealways present in abundance; Vitis rotundifolia is most commonly present, but Toxicodendron radicans, Smilax rotundifolia, Smilaxglauca, and Parthenocissus quinquefolia are usually present in abundance as well. The herbaceous layer may be sparse, particularly ifshrubs and vines are dense, but Chasmanthium laxum may be fairly abundant in this community. Other herbs include Panicumamarum var. amarulum, Eupatorium hyssopifolium, and Elephantopus nudatus. In southern Virginia and North Carolina, Quercusvirginiana and Gelsemium sempervirens may also be present, but Quercus virginiana is never abundant and when present is usuallyrestricted to the understory. Pinus taeda may occur rarely in the Ouachita Mountains and Ozarks of Arkansas where the species isbecoming naturalized, expanding from its native range in the Coastal Plain, where it naturally occurs in low, moist areas (e.g., deep,well-drained soils of floodplains). However, a natural Pinus taeda forest association is not recognized for the Ozark or Ouachitaregion.Related Concepts:• Loblolly Pine: 81 (Eyre 1980) I• Lowland Pine - Oak <strong>Forest</strong> (Foti 1994b) ?• T1A9bII2a. Pinus taeda (Foti et al. 1994) ?• Upland Mixed <strong>Forest</strong> (FNAI 1992a) ?• Upland Mixed <strong>Forest</strong>, Gumbo Loblolly <strong>Forest</strong> subtype (FNAI 1992b) ?Classification Comments: On the Bankhead <strong>National</strong> <strong>Forest</strong> in the Cumberland Plateau of northern Alabama, this alliance includesstreamside terraces that are presumed to have been previously farmed. Associations occurring as plantations are classed in Pinustaeda Planted <strong>Forest</strong> Alliance (A.99).ALLIANCE DISTRIBUTIONRange: This alliance is found in the Cumberland Plateau, Piedmont and Coastal Plains of the southeastern United States, fromDelaware and Maryland south and west to Texas, and in the interior to Tennessee and possibly West Virginia.Subnations: AL, AR, DE, FL, GA, LA, MD, MS, NC, NJ, OK, SC, TN, TX, VATNC Ecoregions: 31:P, 39:C, 40:C, 41:C, 42:P, 43:C, 44:C, 50:C, 52:C, 53:C, 55:?, 56:C, 57:C, 58:C, 59:C, 62:CUSFS Ecoregions: 221D:CC, 221Jb:CCC, 222Cb:CCC, 222Dc:CCC, 222Dd:CCC, 222Eb:CCC, 222Ec:CCC, 222Eg:CCC,231Aa:CCC, 231Ab:CCC, 231Ac:CCC, 231Ad:CCC, 231Ae:CCC, 231Af:CCC, 231Ag:CCC, 231Ah:CCC, 231Ai:CCC,231Aj:CCC, 231Ak:CCC, 231Al:CCC, 231Am:CCC, 231An:CCC, 231Ao:CCP, 231Ba:CCC, 231Bb:CCP, 231Bc:CCP,231Bd:CCC, 231Be:CCP, 231Bf:CCP, 231Bg:CCP, 231Bh:CCP, 231Bi:CCP, 231Bj:CCP, 231Bk:CCP, 231Bl:CC?, 231Ca:CCP,231Cb:CCP, 231Cc:CCP, 231Cd:CCC, 231Ce:CCC, 231Cf:CCC, 231Cg:CCP, 231Da:CCP, 231Dc:CCC, 231De:CC?, 231Ea:CCC,231Eb:CC?, 231Ec:CC?, 231Ed:CC?, 231Ef:CC?, 231Eg:CCP, 231Eh:CCC, 231Ei:CC?, 231Ej:CC?, 231Ek:CCP, 231En:CC?,231Fa:CPP, 231Fb:CP?, 232Ac:CCC, 232Ba:CCC, 232Bb:CC?, 232Bc:CC?, 232Bd:CC?, 232Be:CC?, 232Bg:CCC, 232Bh:CC?,232Bi:CC?, 232Bj:CCC, 232Bk:CC?, 232Bl:CC?, 232Bm:CCC, 232Bn:CC?, 232Bo:CC?, 232Bp:CC?, 232Bq:CCC, 232Br:CCC,232Bt:CC?, 232Bu:CC?, 232Bv:CC?, 232Bx:CC?, 232Bz:CCC, 232Ca:CCC, 232Cb:CCC, 232Cc:CC?, 232Ce:CCC, 232Cf:CC?,232Cg:CC?, 232Ci:CC?, 232Da:CC?, 232Dc:CCC, 232Fa:CC?, 232Fb:CC?, 232Fe:CCC, 255Da:PPP, M221D:??Federal Lands: DOD (Arnold, Fort Benning, Fort Gordon); NPS (Assateague Island, Cape Hatteras, Chickamauga-Chattanooga,Cowpens, Fort Donelson, Guilford Courthouse, Kennesaw Mountain, Kings Mountain, Little River Canyon?, Ninety Six, Shiloh?);TVA (Tellico); USFS (Angelina, Apalachicola, Bankhead, Bienville, Chattahoochee, Conecuh, Croatan, Davy Crockett, Kisatchie,Land Between the Lakes?, Oconee, Sabine NF, Sam Houston, <strong>Sumter</strong>, Talladega, Tuskegee, Uwharrie); USFWS (Chincoteague)ALLIANCE SOURCESReferences: Cain and Shelton 1994, Eyre 1980, FNAI 1992a, FNAI 1992b, Felix et al. 1983, Foti 1994b, Foti et al. 1994, Martin andSmith 1991, Martin and Smith 1993I.A.8.N.B. PINUS ECHINATA FOREST ALLIANCE (A.119)SHORTLEAF PINE FOREST ALLIANCEALLIANCE CONCEPTSummary: This alliance includes forests dominated by Pinus echinata, which on very dry sites may be virtually the only tree speciespresent. This is a wide-ranging alliance; it is currently known from wide areas of the eastern United States from the centralAppalachians south, through the Southern Blue Ridge and Cumberland Plateau and Mountains, possibly extending into the Piedmont,and in the central United States in the Ouachita Mountains and Ozarks, extending south into the Gulf Coastal Plain. Other pinespecies may be present in small amounts; these vary with geography and include Pinus taeda, Pinus virginiana, Pinus pungens, andPinus rigida. Typical hardwood associates include Quercus alba, Quercus falcata, Quercus velutina, Quercus coccinea, Quercusmarilandica, Nyssa sylvatica, Liquidambar styraciflua, Carya alba, and Carya glabra. Understory species vary across the range ofthe alliance, but some common components are Vaccinium arboreum, Vaccinium pallidum, Vaccinium stamineum, SymplocosVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>5
- Page 3: 1 NatureServe is an international o
- Page 11 and 12: types. However, when necessary, mod
- Page 13 and 14: [Association name = floristic nomin
- Page 15 and 16: Generally 5 or fewer occurrences an
- Page 17 and 18: All scientific names for vascular s
- Page 19 and 20: Summary: A short description of the
- Page 21 and 22: Loucks, O. 1996. 100 years after Co
- Page 23 and 24: ALLIANCES BY US NATIONAL VEGETATION
- Page 25: virginiana, and Acer pensylvanicum
- Page 29 and 30: • Appalachian pine-oak forest (Ev
- Page 31 and 32: • Maritime Oak - Holly Forest / W
- Page 33 and 34: ALLIANCE CONCEPTSummary: This allia
- Page 35 and 36: Related Concepts:• Yellow-Poplar:
- Page 37 and 38: virginiana, Sanicula canadensis, De
- Page 39 and 40: Federal Lands: COE (Claiborne Lake)
- Page 41 and 42: ALLIANCE SOURCESReferences: Allard
- Page 43 and 44: Quercus alba and Quercus velutina a
- Page 45 and 46: USFS Ecoregions: 221Hc:CCC, 221He:C
- Page 47 and 48: ALLIANCE CONCEPTSummary: This allia
- Page 49 and 50: USFS Ecoregions: 221Eb:PP?, M221Ab:
- Page 51 and 52: Federal Lands: NPS (Great Smoky Mou
- Page 53 and 54: the associated species. More Appala
- Page 55 and 56: ALLIANCE DISTRIBUTIONRange: This al
- Page 57 and 58: USFS Ecoregions: 221Ha:CCC, 221Hc:C
- Page 59 and 60: elevation environments such as Leio
- Page 61 and 62: CULTIVATED FORESTEASTERN WHITE PINE
- Page 63 and 64: Federal Lands: DOD (Arnold, Fort Be
- Page 65 and 66: does not occur on wet soils. It occ
- Page 67 and 68: Liriodendron tulipifera, although o
- Page 69 and 70: ELEMENT SOURCESReferences: Allard 1
- Page 71 and 72: ELEMENT SOURCESReferences: Allard 1
- Page 73 and 74: Similar Associations:• Pinus echi
- Page 75 and 76: • Pinus echinata - Quercus stella
- Page 77 and 78:
this association occurs in edaphica
- Page 79 and 80:
Vegetation: This forest typically h
- Page 81 and 82:
strobus can have high coverage and
- Page 83 and 84:
CONSERVATION RANKING & RARE SPECIES
- Page 85 and 86:
• Quercus alba - Carya alba / Euo
- Page 87 and 88:
muehlenbergii. In addition, Acer ba
- Page 89 and 90:
WHITE OAK - MOCKERNUT HICKORY / AME
- Page 91 and 92:
CONSERVATION RANKING & RARE SPECIES
- Page 93 and 94:
Vegetation: This association encomp
- Page 95 and 96:
ELEMENT DISTRIBUTIONRange: This com
- Page 97 and 98:
• Quercus prinus - Carya spp. - Q
- Page 99 and 100:
catesbaei, Desmodium nudiflorum, Eu
- Page 101 and 102:
ELEMENT SOURCESReferences: Allard 1
- Page 103 and 104:
High-ranked species: No information
- Page 105 and 106:
Dynamics: No informationSimilar Ass
- Page 107 and 108:
protrusa, Grammitis nimbata (= Micr
- Page 109 and 110:
ELEMENT DISTRIBUTIONRange: This for
- Page 111 and 112:
COMMON WATER-WILLOW HERBACEOUS VEGE
- Page 113 and 114:
• Betula nigra - Platanus occiden
- Page 115 and 116:
Formation: Temporarily flooded temp
- Page 117 and 118:
epeated flooding, this community ma
- Page 119 and 120:
USFS Ecoregions: 221Ba:CCC, 221Ha:C
- Page 121 and 122:
Classification Comments: On the Ban
- Page 123 and 124:
Classification Comments: This veget
- Page 125 and 126:
BIBLIOGRAPHYALNHP [Alabama Natural
- Page 127 and 128:
DeYoung, H. R. 1979. The white pine
- Page 129 and 130:
Gettman, R. W. 1974. A floristic su
- Page 131 and 132:
Lea, C. 2002a. Vegetation classific
- Page 133 and 134:
Pittman, Dr. Albert. Personal commu
- Page 135 and 136:
Tobe, J. D., J. E. Fairey, III, and