procumbens) <strong>Forest</strong> (CEGL006271). Appalachian shale forests and woodlands with Pinus virginiana occur on steep, shaley slopesand have a stunted canopies and sparse herb and shrub strata, characterized by species adapted to shaley substrates. These shalecommunities are classed in Pinus virginiana - Quercus (coccinea, prinus) <strong>Forest</strong> Alliance (A.408) and Pinus (rigida, pungens,virginiana) - Quercus prinus Woodland Alliance (A.677).CONSERVATION RANKING & RARE SPECIESGRank: G4? (2001-2-11): This xeric evergreen forest community will be maintained on sites where local soil conditions,topographic extremes, or occasional fire function to retard hardwood invasion. Infestations of southern pine beetle (Dendroctonusfrontalis) can cause mortality of canopy trees. Examples affected by southern pine beetle in the Great Smoky Mountains can have upto 80-90% standing dead pine.High-ranked species: Buckleya distichophylla (G2), Penstemon deamii (G1), Thermopsis villosa (G3?), Vaccinium hirsutum (G3)ELEMENT DISTRIBUTIONRange: This community occurs primarily in the Appalachian region of the United States, ranging from central Pennsylvania, southand west through the Ridge and Valley, Blue Ridge, and Cumberland Plateau to northern Georgia and Alabama, extending westwardto scattered areas in the Interior Low Plateau and eastward into the upper Piedmont. It is reported from the states of Georgia, NorthCarolina, South Carolina, Tennessee, Kentucky, Pennsylvania, Indiana, Ohio, and is probably in Maryland, Virginia, and WestVirginia.Subnations: AL, GA, IN, KY, MD?, NC, OH, PA, SC, TN, VA?, WVTNC Ecoregions: 43:C, 44:C, 49:C, 50:C, 51:C, 52:C, 59:C, 61:PUSFS Ecoregions: 221Ea:CC?, 221Eb:CCC, 221Ec:CCC, 221Ed:CCP, 221Ef:CCC, 221Eg:CCC, 221Ha:CCC, 221Hb:CCC,221Hc:CCP, 221He:CCC, 221Ja:CCC, 221Jb:CCC, 222Da:CCC, 222Dc:CCC, 222Dg:CCC, 222Dj:CCC, 222Eg:CCC, 222Ej:CCC,222El:CCC, 222En:CCC, 222Eo:CCC, 222Fd:CCC, 222Ff:CCC, 231Aa:CCC, 231Ab:CCC, 231Ae:CCC, 231Bc:CCC, 231Cd:CCC,231Da:CCC, 231Dc:CCC, M221Aa:CCP, M221Ab:CCC, M221Ac:CCC, M221Bd:CCP, M221Be:CCP, M221Cd:CCC,M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Blue Ridge Parkway?, Chickamauga-Chattanooga, Great Smoky Mountains, Kennesaw Mountain, KingsMountain, Little River Canyon?, Mammoth Cave); USFS (Bankhead, Chattahoochee, Cherokee, Daniel Boone, Land Between theLakes?, Nantahala, Pisgah, <strong>Sumter</strong>, Talladega)ELEMENT SOURCESReferences: Allard 1990, Ambrose 1990a, Barden 1977, Burns and Honkala 1990a, CAP pers. comm. 1998, Cooper 1963, Core1966, Evans 1991, Eyre 1980, Fike 1999, Gettman 1974, Homoya pers. comm., Malter 1977, <strong>NatureServe</strong> Ecology - SoutheasternU.S. unpubl. data, Nelson 1986, Patterson et al. 1999, Peet et al. unpubl. data 2002, Pyne 1994, Racine 1966, Rawinski 1992,Schafale 1998b, Schafale and Weakley 1990, Southeastern Ecology Working Group n.d., Walton et al. 1997, Whittaker 1956VIRGINIA PINE SUCCESSIONAL FORESTELEMENT IDENTIFIERSNVC association: Pinus virginiana Successional <strong>Forest</strong>Database Code: CEGL002591Formation: Rounded-crowned temperate or subpolar needle-leaved evergreen forest (I.A.8.N.b)Alliance: Pinus virginiana <strong>Forest</strong> Alliance (A.131)ELEMENT CONCEPTSummary: This community occurs in areas where canopy removal has created dry, open conditions and bare mineral soil, allowingfor the establishment of Pinus virginiana. These habitats include old fields, old pastures, clearcuts, and burned or eroded areas. Thisforest typically has a very dense canopy of Pinus virginiana and little understory vegetation. The dense canopy may also includeadmixtures of other Pinus species (e.g., Pinus taeda, Pinus echinata) or other early successional deciduous trees (e.g., Acer rubrum,Liquidambar styraciflua, Liriodendron tulipifera). Associated woody and herbaceous species vary with geography but are typicallyruderal or exotic species. Shrub and herb layers are frequently very sparse. Stands are short-lived, generally less than 75 years.Environment: This community occurs in areas where canopy removal has created open conditions and bare mineral soil, allowingfor the establishment of Pinus virginiana. These conditions can include old fields, old pastures, clearcuts, and burned or eroded areas.In the Ridge and Valley of Tennessee, northeastern Monroe County, early successional forests with Pinus virginiana dominance werefound on low slopes in areas that were cleared for agriculture prior to the 1970s, when Tellico Lake was created (Andreu and Tukman1995). In the Central Appalachians, this vegetation occurs where soft shales have been farmed (primarily in valleys), resulting instands with nothing but successional species in the understory. Soils underlying these communities are of two general types, i.e., thosederived in residuum from calcareous shale and calcareous sandstone of the Middle Ordovician and those of some other origin. Seriesof the former type include Dandridge (Lithic Ruptic-Alfic Eutrochrepts), Tellico (Typic Rhododults), and Steekee (Ruptic-UlticDystrochrepts). Other soil series that this forest type may occur on include Litz, Dewey, Alcoa, Bland, Etowah, Lobdell and Neubert.All of these soils are well-drained and range in pH from moderate acid to very strongly acidic.Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>56
Vegetation: This forest typically has a very dense canopy of Pinus virginiana and little understory vegetation. Pinus taeda or Pinusechinata may co-occur with Pinus virginiana in the canopy. The canopy can also have significant admixtures of early successionaldeciduous trees (e.g., Acer rubrum, Liquidambar styraciflua, Liriodendron tulipifera). Associated woody and herbaceous species varywith geography but are typically ruderal or exotic species. Shrub and herb strata are absent to sparse in coverage. In eastern Tennesseethe subcanopy may contain Acer saccharum and Cornus florida; other associated species may include Cercis canadensis,Parthenocissus quinquefolia, Lonicera japonica, and Microstegium vimineum (Andreu and Tukman 1995). In the CentralAppalachians, associates include Pinus taeda, Pinus echinata, and Pinus rigida. The dense ericaceous shrub stratum containsVaccinium spp., Gaylussacia spp., Kalmia latifolia, and Rhododendron spp.Dynamics: This is an early successional forest type. Damage from ice storms was the main disturbance observed in these stands inthe Tellico Pilot Project study area. In addition, fire and insect infestation are likely damaging agents.Similar Associations:• Pinus virginiana - Juniperus virginiana var. virginiana - Ulmus alata <strong>Forest</strong> (CEGL007121) -- on more calcareous orcircumneutral substrates.Related Concepts:• IA7c. Xeric Virginia Pine Ridge <strong>Forest</strong> (Allard 1990) B• Unclassified Old-Field Successional <strong>Forest</strong> (Fleming and Moorhead 2000) ?• Virginia Pine - Oak: 78 (Eyre 1980) B• Virginia Pine, RV (Pyne 1994) B• Virginia Pine: 79 (Eyre 1980) B Xeric Pine <strong>Forest</strong> (Ambrose 1990a) BClassification Comments: Early successional Pinus virginiana vegetation occurring over calcareous substrates is classed in Pinusvirginiana - Juniperus virginiana var. virginiana - Ulmus alata <strong>Forest</strong> (CEGL007121) and has species indicative of calcareoussubstrates.CONSERVATION RANKING & RARE SPECIESGRank: GNA (ruderal) (2000-6-13): This forest represents early successional vegetation and is thus not of conservation concern.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This successional community is possible in the Piedmont from Pennsylvania south to Alabama, and ranges west into theAppalachians, Ridge and Valley, the Cumberland Plateau, and in scattered locales of the Interior Low Plateau.Subnations: AL, GA, IN, KY, MD, NC, NJ, PA, SC, TN, VA, WVTNC Ecoregions: 50:C, 51:C, 52:C, 58:P, 59:C, 61:CUSFS Ecoregions: 221D:CC, 221Ha:CCC, 221Hb:CCC, 221Hc:CCC, 221He:CCC, 221J:CC, 222Ej:CCC, 222En:CCC,222Eo:CCC, 231Cd:CCC, M221Aa:CCC, M221Ab:CCC, M221Ac:CCC, M221Ca:CCP, M221Cb:CCP, M221Cc:CCP,M221Cd:CCC, M221Ce:CCP, M221Da:CCC, M221Db:CCC, M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Great Smoky Mountains, Little River Canyon?, Shiloh); TVA (Tellico); USFS (Bankhead, Chattahoochee,Cherokee, Daniel Boone, George Washington, Jefferson, <strong>Sumter</strong>, Uwharrie?)ELEMENT SOURCESReferences: Allard 1990, Ambrose 1990a, Andreu and Tukman 1995, Eyre 1980, Fike 1999, Fleming and Coulling 2001, Flemingand Moorhead 2000, Nelson 1986, Patterson et al. 1999, Pyne 1994, Southeastern Ecology Working Group n.d.SOUTHERN APPALACHIAN MONTANE CLIFF AND TALUSMOUNTAIN SPLEENWORT - ROCK ALUMROOT FELSIC CLIFF SPARSE VEGETATIONELEMENT IDENTIFIERSNVC association: Asplenium montanum - Heuchera villosa Felsic Cliff Sparse VegetationDatabase Code: CEGL004980Formation: Cliffs with sparse vascular vegetation (VII.A.1.N.a)Alliance: Asplenium montanum Sparsely Vegetated Alliance (A.1831)ELEMENT CONCEPTSummary: This community occurs in the Blue Ridge and upper Piedmont of Georgia, North Carolina, South Carolina, Tennessee,and Virginia. It includes vertical rock faces associated with felsic, metamorphic and igneous geologies. This community generally haslittle vegetative cover, often with 90% of the rock surface unvegetated. Mosses (e.g., Thuidium spp., Fissidens spp., Campylium sp.,Bryoandersonia sp., Plagiomnium sp.) and lichens can have moderate coverage, and vascular plants occur on ledges and rooted incracks. Asplenium montanum and Heuchera villosa are characteristic components. Other typical species include Agrostis perennans,Arisaema triphyllum, Aristolochia macrophylla, Asplenium trichomanes, Eurybia divaricata (= Aster divaricatus), Cystopterisprotrusa, Dryopteris marginalis, Hydrangea arborescens, Parthenocissus quinquefolia, Polypodium appalachianum, and RubusVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>57
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1 NatureServe is an international o
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types. However, when necessary, mod
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[Association name = floristic nomin
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Generally 5 or fewer occurrences an
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Summary: A short description of the
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Loucks, O. 1996. 100 years after Co
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ALLIANCES BY US NATIONAL VEGETATION
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virginiana, and Acer pensylvanicum
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- Page 41 and 42: ALLIANCE SOURCESReferences: Allard
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- Page 65 and 66: does not occur on wet soils. It occ
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- Page 71 and 72: ELEMENT SOURCESReferences: Allard 1
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- Page 109 and 110: ELEMENT DISTRIBUTIONRange: This for
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- Page 113 and 114: • Betula nigra - Platanus occiden
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- Page 117 and 118: epeated flooding, this community ma
- Page 119 and 120: USFS Ecoregions: 221Ba:CCC, 221Ha:C
- Page 121 and 122: Classification Comments: On the Ban
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- Page 125 and 126: BIBLIOGRAPHYALNHP [Alabama Natural
- Page 127 and 128: DeYoung, H. R. 1979. The white pine
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Gettman, R. W. 1974. A floristic su
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Lea, C. 2002a. Vegetation classific
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Pittman, Dr. Albert. Personal commu
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Tobe, J. D., J. E. Fairey, III, and