examples of this community. No effort has been made to subdivide this type into northern and southern, or Coastal Plain and Interiorvariants, although there are undoubtedly some floristic differences between these extremes, at least in the lower strata.Dynamics: <strong>Forest</strong>s in this alliance occur on riverfronts in areas with repeated, frequent, natural disturbance in the form of flooding.Occurrences of this type may have a rather high percentage of standing dead trees, including remnant snags from earlier successionalcommunities. A layer of sand is often deposited at the surface of the soil when streams overflow their banks during flash floods. Littleor no clay is present in the upper strata of soils which support stands of this alliance. Flooding is seasonal and/or occasional andshallow, but never prolonged. Most of this short-duration flooding takes place in early spring.This is a Zone IV community. Zone IV communities experience seasonal inundation or groundwater saturation for approximately12.5-25% of the year, usually during spring and early summer with a frequency of 51-100%.This community often succeeds riverfront forests dominated by Salix spp. and Populus spp. It probably succeeds to forests dominatedby Celtis spp., Ulmus spp., Fraxinus pennsylvanica, and Liquidambar styraciflua.Similar Associations:• Betula nigra - Platanus occidentalis <strong>Forest</strong> (CEGL002086)Related Concepts:• Brownwater Levee <strong>Forest</strong> (Birch-Sycamore Successional Phase) (Schafale 2000) ?• Eutrophic Seasonally Flooded <strong>Forest</strong> (Rawinski 1992) ?• IIA7b. River Birch - Sycamore Riverfront <strong>Forest</strong> (Allard 1990) B• Mountain Stream Floodplain <strong>Forest</strong> (Ambrose 1990a) ?• River Birch - Sycamore (72) (USFS 1988) ?• River Birch - Sycamore Riverfront <strong>Forest</strong> (Oberholster 1993) B• River Birch - Sycamore: 61 (Eyre 1980) ?• River Birch-Sycamore Riverfront <strong>Forest</strong> (Pyne 1994) ?Classification Comments: No effort has been made to subdivide this type into northern and southern, or Coastal Plain and Interiorvariants, although there are undoubtedly some floristic differences between these extremes, at least in the lower strata. This includesthe 'Sycamore - River Birch - Silver Maple' forests of the Pearl River in Louisiana mentioned by K. Ribbeck. Additional species fromthe shrub and herb strata need to be added to make more apparent the differences between this association and Betula nigra - Platanusoccidentalis <strong>Forest</strong> (CEGL002086), which is a more northern forest. In Kentucky, this association (CEGL007312) would be found onthe southern part of the Daniel Boone <strong>National</strong> <strong>Forest</strong>, in contrast to CEGL002086, which is more northern.CONSERVATION RANKING & RARE SPECIESGRank: G4G5 (2001-9-19): This is a widespread, variable, dynamic association. The principal threats to it come from hydrologicalterations, stands do not typically contain valuable timber species.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This riverfront association ranges from Virginia to southern Georgia in the Piedmont and Coastal Plain (and into the southernand lower-elevation parts of the Southern Blue Ridge, excluding the highest elevations of the mountains) and west to eastern Texas,and possibly the Cumberland Plateau of northern Alabama.Subnations: AL, GA, KY, LA, MS, NC, SC, TN, TX, VATNC Ecoregions: 32:P, 40:P, 41:C, 42:C, 43:C, 44:C, 50:C, 51:C, 52:C, 53:C, 56:C, 57:C, 58:?USFS Ecoregions: 221Hc:CCC, 222Cf:CCC, 222Cg:CCC, 222E:CC, 231Cd:CPP, 231E:CC, 231Ga:CCC, 231Gb:CCC,231Gc:CCC, 232B:CC, 232C:CC, 232D:CC, 232F:CC, 234A:CC, 255:C, M221Cd:CCC, M221Dc:CCC, M231Aa:CCC,M231Ab:CCC, M231Ac:CCC, M231Ad:CCCFederal Lands: DOD (Fort Benning); NPS (Shiloh); USFS (Bankhead?, Chattahoochee, Daniel Boone, Kisatchie, <strong>Sumter</strong>); USFWS(Little River)ELEMENT SOURCESReferences: Allard 1990, Ambrose 1990a, Burns and Honkala 1990b, Campbell 1988, Campbell 1989a, Clark and Benforado 1981,Dickson and Segelquist 1978, Duever and Brinson 1984b, Evans 1991, Eyre 1980, FNAI 1992b, Faulkner and Patrick n.d., Fleminget al. 2001, Flinchum 1977, Gettman 1974, Hoagland 1997, Kartesz 1999, Klimas et al. 1981, McWilliams and Rosson 1990,<strong>NatureServe</strong> Ecology - Southeastern U.S. unpubl. data, Nelson 1986, Oberholster 1993, Palmer-Ball et al. 1988, Peet et al. unpubl.data 2002, Putnam 1951, Putnam et al. 1960, Pyne 1994, Rawinski 1992, Schafale 2000, Schafale and Weakley 1990, Smith 1996a,Southeastern Ecology Working Group n.d., USFS 1988, Wharton 1978, Wharton et al. 1982, Wieland 1994b, Wieland 2000bVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>88
COMMON WATER-WILLOW HERBACEOUS VEGETATIONELEMENT IDENTIFIERSNVC association: Justicia americana Herbaceous VegetationDatabase Code: CEGL004286Formation: Temporarily flooded temperate perennial forb vegetation (V.B.2.N.d)Alliance: Justicia americana Temporarily Flooded Herbaceous Alliance (A.1657)SOUTH-CENTRAL INTERIOR SMALL STREAM AND RIPARIANELEMENT CONCEPTSummary: This association is found primarily in the Piedmont, Cumberland Plateau, Interior Low Plateau, Ozarks, OuachitaMountains, and adjacent provinces. Stands occur on the shoals or bars of rocky streams and riverbeds. It provides habitat in someportions of its range for globally rare dragonflies and herbs. Justicia americana is the characteristic dominant. Other herbaceousspecies that may be present include Diodia teres, Gratiola brevifolia, Leersia sp., Lemna minor, Orontium aquaticum, Podostemumceratophyllum, Scirpus sp., Saururus cernuus, and Xyris difformis var. difformis. A sparse canopy layer, which can include Carpinuscaroliniana ssp. caroliniana, Salix interior, Fagus grandifolia, and Fraxinus pennsylvanica among other species, may be present.Environment: This association occurs on the shoals or bars of rocky streams and riverbeds, or gravelly sands.Vegetation: Justicia americana is the characteristic dominant. Other herbaceous species that may be present include Diodia teres,Gratiola brevifolia, Leersia sp., Lemna minor, Orontium aquaticum, Podostemum ceratophyllum, Saururus cernuus, and Xyrisdifformis var. difformis. In Ohio, Justicia usually grows in nearly pure patches, so that few other species are associated with it. Bidensspp., Cuscuta gronovii, Mimulus ringens, Polygonum spp., Rumex spp., and Salix interior can occur (Anderson 1982). A sparsecanopy layer may be present, which can include Carpinus caroliniana, Fagus grandifolia, and Fraxinus pennsylvanica, among others.In the Cumberland Plateau of Alabama, Justicia americana is present in dense patches with some interspersion of other speciesincluding Pilea pumila, Boehmeria cylindrica, Eclipta prostrata (= Eclipta alba), Juncus coriaceus, Mikania scandens, Ludwigiapalustris, Leersia sp. and Bidens sp. Schmalzer and DeSelm (1982) discuss Orontium aquaticum growing along streambanks or inshallow riffles "along or with" Justicia americana in the Obed River in the Cumberland Plateau of Tennessee.Dynamics: Stands in some situations may be obliterated by ongoing river channeling. Anderson (1982) describes some of the lifehistorycharacteristics of Justicia americana that allow it to persist in river channels.Similar Associations:Related Concepts:• Aquatic Types (Schmalzer and DeSelm 1982) B• IIE3a. Riverside Shoal and Stream Bar Complex (Allard 1990) B• Rocky Bar and Shore (Water Willow Subtype) (Schafale 1998b) ?Classification Comments: This type, in Ohio, often forms pure patches, but consistent identification may require a simple cutoffrule, such as at least 50% cover of Justicia (Anderson 1982). However, Anderson (1996) no longer recognizes this type.CONSERVATION RANKING & RARE SPECIESGRank: G4G5 (1997-9-12): No informationHigh-ranked species: No informationELEMENT DISTRIBUTIONRange: This type is found primarily in the Piedmont, Interior Low Plateau, Cumberland Plateau, Ozarks, Ouachita Mountains, andadjacent provinces. It ranges from Alabama, Georgia and the Carolinas west to Arkansas and Oklahoma and north to Ohio,Pennsylvania, and Delaware.Subnations: AL, AR, GA, KY, MD?, NC, OH, OK, PA, SC?, TN, VA?, WVTNC Ecoregions: 38:C, 39:C, 43:C, 44:C, 45:C, 48:C, 49:C, 50:C, 51:C, 52:C, 59:C, 60:C, 61:CUSFS Ecoregions: 212Fa:CCP, 212Fb:CCC, 212Ga:CCP, 212Gb:CCP, 221Am:CCP, 221Ba:CCC, 221Bd:CCC, 221Da:CCC,221Db:CCC, 221Ec:CCC, 221Ed:CCP, 221Ef:CCP, 221Ha:CCC, 221Hb:CCC, 221Hc:CCC, 221He:CCC, 222Eb:CCC, 222Ej:CCP,222En:CCC, 222Eo:CCC, 222Ha:CCC, 231Af:CCC, 231B:CC, 231Cd:CCC, 231D:CC, M221Aa:CCC, M221Ac:CCC,M221Ad:CCC, M221Bb:CCC, M221Bd:CCC, M221Be:CCC, M221Bf:CCC, M221Cd:CCC, M221Da:CCC, M221Dc:CCC,M222A:CC, M231A:CCFederal Lands: NPS (Little River Canyon?, Natchez Trace, Stones River); USFS (Bankhead, Cherokee, Daniel Boone, Oconee?,Ouachita, Ozark, Pisgah, <strong>Sumter</strong>?, Uwharrie)ELEMENT SOURCESReferences: Allard 1990, Anderson 1982, Anderson 1996, Fike 1999, Fleming et al. 2001, Hoagland 1997, Hoagland 2000, Major etal. 1999, McCoy 1958, Nelson 1986, Palmer-Ball et al. 1988, Peet et al. unpubl. data 2002, Penfound 1953, Schafale 1998b, Schafaleand Weakley 1990, Schmalzer and DeSelm 1982, Southeastern Ecology Working Group n.d.Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>89
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1 NatureServe is an international o
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types. However, when necessary, mod
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[Association name = floristic nomin
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Generally 5 or fewer occurrences an
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All scientific names for vascular s
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Summary: A short description of the
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Loucks, O. 1996. 100 years after Co
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ALLIANCES BY US NATIONAL VEGETATION
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virginiana, and Acer pensylvanicum
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virginiana, Juniperus virginiana va
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• Appalachian pine-oak forest (Ev
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• Maritime Oak - Holly Forest / W
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ALLIANCE CONCEPTSummary: This allia
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Related Concepts:• Yellow-Poplar:
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virginiana, Sanicula canadensis, De
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Federal Lands: COE (Claiborne Lake)
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ALLIANCE SOURCESReferences: Allard
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Quercus alba and Quercus velutina a
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USFS Ecoregions: 221Hc:CCC, 221He:C
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ALLIANCE CONCEPTSummary: This allia
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USFS Ecoregions: 221Eb:PP?, M221Ab:
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Federal Lands: NPS (Great Smoky Mou
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the associated species. More Appala
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ALLIANCE DISTRIBUTIONRange: This al
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USFS Ecoregions: 221Ha:CCC, 221Hc:C
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- Page 125 and 126: BIBLIOGRAPHYALNHP [Alabama Natural
- Page 127 and 128: DeYoung, H. R. 1979. The white pine
- Page 129 and 130: Gettman, R. W. 1974. A floristic su
- Page 131 and 132: Lea, C. 2002a. Vegetation classific
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- Page 135 and 136: Tobe, J. D., J. E. Fairey, III, and