canadensis, Adiantum pedatum, Polystichum acrostichoides, Ageratina altissima var. roanensis, Hepatica nobilis var. acuta, Asarumcanadense, Stellaria pubera, Tiarella cordifolia, Clintonia umbellulata, Sedum ternatum, Mitella diphylla, Osmorhiza claytonii,Dryopteris intermedia, Arisaema triphyllum, Cystopteris protrusa, Trillium grandiflorum, Viola canadensis, Dicentra canadensis,Dicentra cucullaria, Hydrophyllum canadense, Hydrophyllum virginianum, Phacelia bipinnatifida, Phacelia fimbriata, Delphiniumtricorne, Carex austrocaroliniana, Carex manhartii, Carex plantaginea, and Carex platyphylla. These forests mainly occur onprotected, mesic, low to moderate elevation (2000-4500 feet, 610-1370 m) sites, primarily broad coves and lower slopes. <strong>Forest</strong>s inthis alliance are known from the Southern Blue Ridge of North Carolina, South Carolina, Georgia, Tennessee, Virginia, and theCumberland Mountains of Kentucky. More information is needed to characterize forests provisionally assigned to this alliance thatoccur in the Allegheny Plateau of West Virginia and Ohio and in Indiana.Related Concepts:• Appalachian mesophytic forest (Evans 1991) I• Beech - Sugar Maple: 60 (Eyre 1980) I• IA5a. Southern Appalachian Mesophytic Cove <strong>Forest</strong> (Allard 1990) ?• Mesic Central <strong>Forest</strong> (Smith 1991) I• Mixed Mesophytic BR (Pyne 1994) ?• Mixed mesophytic forest (Fike 1999) ?• Rich Cove <strong>Forest</strong> (Schafale and Weakley 1990) ?• Yellow-Poplar: 57 (Eyre 1980) IClassification Comments: NoneALLIANCE DISTRIBUTIONRange: This alliance is found in southern Indiana, southern Ohio, West Virginia, Virginia (?), Georgia, Kentucky, North Carolina,South Carolina, and Tennessee. <strong>Forest</strong>s in this alliance are known from the Southern Blue Ridge of North Carolina, South Carolina,Georgia, Tennessee, Virginia, and the Cumberland Mountains of Kentucky. More information is needed to characterize forestsprovisionally assigned to this alliance that occur in the Allegheny Plateau of West Virginia and Ohio and in Indiana.Subnations: GA, IN, KY, NC, OH, PA, SC, TN, VA, WVTNC Ecoregions: 44:C, 49:C, 50:C, 51:C, 52:C, 59:CUSFS Ecoregions: 221Ea:CCC, 221Eb:CCC, 221Ec:CCC, 221Ed:CCC, 221Ee:CCC, 221Ef:CCC, 221Eg:CCC, 221Ha:CPP,221Hb:CPP, 221Hc:CPP, 221He:CPP, 222E:CC, 222F:CC, 231Ab:CCC, 231Ad:CCC, M221Aa:CCC, M221Ab:CCC,M221Bb:CCC, M221Bd:CCC, M221Be:CCC, M221Cc:CC?, M221Cd:CCC, M221Ce:CCC, M221Da:CCC, M221Db:CCC,M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Blue Ridge Parkway, Great Smoky Mountains); USFS (Chattahoochee, Cherokee, Daniel Boone, GeorgeWashington, Jefferson, Nantahala, Pisgah, <strong>Sumter</strong>)ALLIANCE SOURCESReferences: Allard 1990, Boufford and Wood 1977, Chapman 1957, Cooper 1963, Cooper and Hardin 1970, Dellinger unpubl. data1992, DuMond 1970, Evans 1991, Eyre 1980, Faber-Langendoen et al. 1996, Fike 1999, Golden 1974, Govus 1982, Greenlee 1974,Malter 1977, McLeod 1988, Newell et al. 1997, Patterson 1994, Pyne 1994, Rodgers and Shake 1965, Schafale and Weakley 1990,Smith 1991, Thomas 1966, Tobe et al. 1992, Tucker 1973, Weakley et al. 1979, Whigham 1969, Whittaker 1956I.B.2.N.A. LIRIODENDRON TULIPIFERA FOREST ALLIANCE (A.236)TULIPTREE FOREST ALLIANCEALLIANCE CONCEPTSummary: This alliance includes deciduous forests dominated by Liriodendron tulipifera, primarily in areas which were onceclearcut, old fields, or cleared by fire or other natural disturbances. These non-wetland forests are also found along mesic streamterraces and on upland mountain benches. <strong>Forest</strong>s in this alliance are abundant in the central and southern Appalachians, below 915 m(3000 feet) elevation, usually associated with disturbance and on the most productive sites, but also occur in the Coastal Plain,Piedmont, Ridge and Valley, and Cumberland Plateau. This alliance includes pure, often even-aged stands of Liriodendron tulipiferaas well as forests with Liriodendron tulipifera associated with other species favored by canopy openings. Associated species varywith geographic location. Throughout most of the range of this alliance, Acer rubrum, Robinia pseudoacacia, Betula lenta, Acersaccharum, and Acer negundo are common components. In the Piedmont and Coastal Plain, Liquidambar styraciflua is a commonassociate. In the Appalachians, Halesia tetraptera, Tsuga canadensis, Tilia americana var. heterophylla (= Tilia heterophylla),Prunus serotina var. serotina, and Magnolia fraseri can be additional components. In the Ridge and Valley and Cumberland Plateau,additional species include Quercus rubra, Magnolia acuminata, Carya alba, Carya glabra, Pinus virginiana, Sassafras albidum,Pinus strobus, Carpinus caroliniana, Asimina triloba, and Staphylea trifolia. Herbaceous strata are not diverse and, in the southernAppalachians, this feature distinguishes these forests from rich cove forests in Liriodendron tulipifera - Tilia americana var.heterophylla - Aesculus flava - Acer saccharum <strong>Forest</strong> Alliance (A.235). Vines can be abundant including Vitis spp., Smilax spp.,Aristolochia macrophylla, and Parthenocissus quinquefolia. <strong>Forest</strong>s in this alliance occur on middle to lower slopes, sheltered covesand gentle concave slopes, and river terraces over various soils and geologies. Vegetation of this alliance is uncommon in Louisiana.Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>12
Related Concepts:• Yellow-Poplar: 57 (Eyre 1980) IClassification Comments: NoneALLIANCE DISTRIBUTIONRange: This alliance is found in Alabama, Georgia, Kentucky, Louisiana, Mississippi (?), North Carolina, South Carolina,Tennessee, Maryland, Pennsylvania, Virginia, and West Virginia. <strong>Forest</strong>s in this alliance are abundant in the central and southernAppalachians, below 915 m (3000 feet) elevation, but also occur in the Coastal Plain, Piedmont, Ridge and Valley, and CumberlandPlateau.Subnations: AL, GA, KY, LA, MD, MS?, NC, PA, SC, TN, VA, WVTNC Ecoregions: 43:C, 44:C, 50:C, 51:C, 52:C, 53:P, 58:C, 59:CUSFS Ecoregions: 221Ha:CCC, 221Hc:CCC, 221He:CCC, 221Jb:CCC, 222C:CC, 222D:CC, 222Eb:CCC, 222Ed:CCP,222En:CCC, 222Eo:CCC, 231Aa:CCP, 231Ae:CCC, 231Bc:CCC, 231Cd:CCC, 231Dc:CCC, 232B:CC, 232D:CP, 234Ab:CCC,M221Aa:CCC, M221Ab:CCP, M221Ac:CCC, M221Ad:CCC, M221Bb:CCC, M221Da:CCC, M221Db:CCP, M221Dc:CCC,M221Dd:CCCFederal Lands: DOD (Arnold, Fort Benning); NPS (Blue Ridge Parkway, Cowpens, Fort Donelson, Great Smoky Mountains,Guilford Courthouse, Harpers Ferry, Kennesaw Mountain, Kings Mountain, Ninety Six, Rock Creek, Shenandoah, Shiloh); TVA(Tellico); USFS (Apalachicola, Bankhead, Bienville, Chattahoochee, Cherokee, Conecuh, Daniel Boone, De Soto, GeorgeWashington, Jefferson, Nantahala, Oconee?, Pisgah, St. Francis, <strong>Sumter</strong>, Talladega, Tombigbee, Tuskegee)ALLIANCE SOURCESReferences: Andreu and Tukman 1995, Eyre 1980, Gallyoun et al. 1996, Golden 1974, Horn 1980, McGee and Hooper 1970,Phillips and Shure 1990, Schmalzer 1978, Thomas 1966I.B.2.N.A. QUERCUS ALBA - (QUERCUS RUBRA, CARYA SPP.) FOREST ALLIANCE (A.239)WHITE OAK - (NORTHERN RED OAK, HICKORY SPECIES) FOREST ALLIANCEALLIANCE CONCEPTSummary: This alliance is widely distributed in the eastern United States and portions of adjacent Canada and includes dry mesic tomesic upland oak forests dominated by Quercus alba and/or Quercus rubra, with or without Carya species. Stands are 15-25 m tall,with a closed, deciduous canopy. The shrub and herbaceous strata are typically well-developed. Quercus alba usually dominates thestands, either alone or in combination with Quercus rubra (especially on moister sites) and sometimes Quercus velutina (especially ondrier sites). Some associations in this alliance are dominated by Quercus rubra, although Quercus alba is usually also a canopycomponent. Carya species (particularly Carya alba, Carya glabra or Carya ovata) are typically common either in the canopy orsubcanopy. In the southeastern United States, this alliance covers dry-mesic forests of the Piedmont, low Appalachian Mountains, andthe Cumberland and Interior Low Plateau, and mesic oak-hickory forests of the Blue Ridge and the interior highlands of the Ozarksand Ouachita Mountains. Associated species include Carya glabra, Carya ovata, Carya alba, Fraxinus americana, Acer rubrum,Acer leucoderme, Cornus florida, Nyssa sylvatica, Ostrya virginiana, Calycanthus floridus, Pyrularia pubera, Tilia americana var.caroliniana, Oxydendrum arboreum, and others. This alliance is found throughout the midwestern United States on moderately rich,upland sites. Typical associates include Fraxinus americana, Ulmus americana, Tilia americana, Acer saccharum, Acer rubrum, andmore locally, Quercus macrocarpa and Quercus ellipsoidalis.Stands are found on gentle to moderately steep slopes on uplands and on steep valley sides. The soils are moderately deep to deep andvary from silts to clays and loams. The parent material ranges from glaciated till to limestone, shale, sandstone and other bedrocktypes. In the midwestern United States, many stands are succeeding to types dominated by Acer saccharum, Tilia americana, Acerrubrum, and other mesic tree associates. This succession may be delayed by fire and grazing. In the eastern and southeastern UnitedStates, Liriodendron tulipifera, Fraxinus americana, Acer rubrum, and other mesic associates often increase after disturbances, suchas clearcutting or windstorms, especially in the absence of fire.Related Concepts:• Acidic mesophytic forest (Evans 1991) I• Basic Oak - Hickory <strong>Forest</strong> (Nelson 1986) ?• Basic Oak--Hickory <strong>Forest</strong>, Mafic Substrate Variant (Schafale and Weakley 1990) I• Calcareous mesophytic forest (Evans 1991) I• Dry-Mesic Oak--Hickory <strong>Forest</strong> (Schafale and Weakley 1990) ?• IA6j. Interior Calcareous Oak - Hickory <strong>Forest</strong> (Allard 1990) I• Mesic Oak - Hickory <strong>Forest</strong> (Foti 1994b) I• Montane Oak--Hickory <strong>Forest</strong> (Schafale and Weakley 1990) I• Oak - Chestnut - Hickory <strong>Forest</strong> (Ambrose 1990a) I• Oak - Hickory <strong>Forest</strong> (Swain and Kearsley 2001) ?• Oak--Hickory <strong>Forest</strong> (Nelson 1986) IVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>13
- Page 3: 1 NatureServe is an international o
- Page 11 and 12: types. However, when necessary, mod
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- Page 15 and 16: Generally 5 or fewer occurrences an
- Page 17 and 18: All scientific names for vascular s
- Page 19 and 20: Summary: A short description of the
- Page 21 and 22: Loucks, O. 1996. 100 years after Co
- Page 23 and 24: ALLIANCES BY US NATIONAL VEGETATION
- Page 25 and 26: virginiana, and Acer pensylvanicum
- Page 27 and 28: virginiana, Juniperus virginiana va
- Page 29 and 30: • Appalachian pine-oak forest (Ev
- Page 31 and 32: • Maritime Oak - Holly Forest / W
- Page 33: ALLIANCE CONCEPTSummary: This allia
- Page 37 and 38: virginiana, Sanicula canadensis, De
- Page 39 and 40: Federal Lands: COE (Claiborne Lake)
- Page 41 and 42: ALLIANCE SOURCESReferences: Allard
- Page 43 and 44: Quercus alba and Quercus velutina a
- Page 45 and 46: USFS Ecoregions: 221Hc:CCC, 221He:C
- Page 47 and 48: ALLIANCE CONCEPTSummary: This allia
- Page 49 and 50: USFS Ecoregions: 221Eb:PP?, M221Ab:
- Page 51 and 52: Federal Lands: NPS (Great Smoky Mou
- Page 53 and 54: the associated species. More Appala
- Page 55 and 56: ALLIANCE DISTRIBUTIONRange: This al
- Page 57 and 58: USFS Ecoregions: 221Ha:CCC, 221Hc:C
- Page 59 and 60: elevation environments such as Leio
- Page 61 and 62: CULTIVATED FORESTEASTERN WHITE PINE
- Page 63 and 64: Federal Lands: DOD (Arnold, Fort Be
- Page 65 and 66: does not occur on wet soils. It occ
- Page 67 and 68: Liriodendron tulipifera, although o
- Page 69 and 70: ELEMENT SOURCESReferences: Allard 1
- Page 71 and 72: ELEMENT SOURCESReferences: Allard 1
- Page 73 and 74: Similar Associations:• Pinus echi
- Page 75 and 76: • Pinus echinata - Quercus stella
- Page 77 and 78: this association occurs in edaphica
- Page 79 and 80: Vegetation: This forest typically h
- Page 81 and 82: strobus can have high coverage and
- Page 83 and 84: CONSERVATION RANKING & RARE SPECIES
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• Quercus alba - Carya alba / Euo
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muehlenbergii. In addition, Acer ba
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WHITE OAK - MOCKERNUT HICKORY / AME
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CONSERVATION RANKING & RARE SPECIES
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Vegetation: This association encomp
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ELEMENT DISTRIBUTIONRange: This com
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• Quercus prinus - Carya spp. - Q
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catesbaei, Desmodium nudiflorum, Eu
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ELEMENT SOURCESReferences: Allard 1
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High-ranked species: No information
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Dynamics: No informationSimilar Ass
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protrusa, Grammitis nimbata (= Micr
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ELEMENT DISTRIBUTIONRange: This for
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COMMON WATER-WILLOW HERBACEOUS VEGE
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• Betula nigra - Platanus occiden
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Formation: Temporarily flooded temp
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epeated flooding, this community ma
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USFS Ecoregions: 221Ba:CCC, 221Ha:C
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Classification Comments: On the Ban
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Classification Comments: This veget
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BIBLIOGRAPHYALNHP [Alabama Natural
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DeYoung, H. R. 1979. The white pine
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Gettman, R. W. 1974. A floristic su
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Lea, C. 2002a. Vegetation classific
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Pittman, Dr. Albert. Personal commu
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Tobe, J. D., J. E. Fairey, III, and