WETLANDS, VEGETATEDATLANTIC COASTAL PLAIN SMALL BROWNWATER RIVER FLOODPLAIN FORESTEASTERN COTTONWOOD - BLACK WILLOW / CLIMBING HEMPVINE FORESTELEMENT IDENTIFIERSNVC association: Populus deltoides - Salix nigra / Mikania scandens <strong>Forest</strong>Database Code: CEGL007346Formation: Temporarily flooded cold-deciduous forest (I.B.2.N.d)Alliance: Populus deltoides Temporarily Flooded <strong>Forest</strong> Alliance (A.290)ELEMENT CONCEPTSummary: This forest, dominated by Populus deltoides and Salix nigra, occurs in the southeastern Coastal Plain along the fronts andbanks of large rivers and on islands where sediment accretes. In addition to the above-listed species, Acer rubrum, Fraxinuspennsylvanica, Salix caroliniana, Betula nigra, Planera aquatica, Ulmus americana, Liquidambar styraciflua, Morus rubra, Juglansnigra, and Acer saccharinum all may be present within their ranges. Ilex opaca var. opaca, Carpinus caroliniana, Lindera benzoin,Cornus drummondii, and <strong>Forest</strong>iera acuminata may be present in the subcanopy. Depending on the maturity of the occurrence,physiognomy varies. The herb layer is sparse to dense. Distribution is not known. The western limit of distribution also is not known.Work needs to be done to determine more precisely the range of this community and to determine whether it needs to be separatedinto less widespread types. The ranges of the dominant species in all strata do not preclude its wide distribution.Environment: This forest occurs in the southeastern Coastal Plain along the fronts and banks of large rivers and on islands wheresediment accretes. This community is found on the fronts or banks of major rivers which are better-drained than areas farther from thewatercourse. It can also be found on relatively newly accreted soil at the ends of islands and stream bars. Soils are coarser-texturedthan those of most other bottomland communities because the heavier soil particles drop out of flood waters first; silts and clays staysuspended longer and are deposited farther from the watercourse. This community may establish itself in either Zone III or Zone IV.If it occurs in Zone III, soil accretion will move the community into Zone IV. Zone III communities experience semipermanentlyinundated or saturated soil from surface or groundwater during >25% of the growing season; flooding is typical during winter andspring with a 51-100% frequency, but the soil is not saturated approximately 60% of the year. Zone IV communities experienceseasonal inundation or groundwater saturation for approximately 12.5-25% of the year, usually during spring and early summer with afrequency of 51-100%.Vegetation: Stands of this forest are dominated by Populus deltoides and Salix nigra. These two species combined contribute at least75% of the canopy cover with each contributing 25-75% of the total and a variety of other bottomland species contributing the rest. Inaddition to the above-listed species, Acer rubrum, Fraxinus pennsylvanica, Salix caroliniana, Betula nigra, Planera aquatica, Ulmusamericana, Liquidambar styraciflua, Morus rubra, Juglans nigra, and Acer saccharinum all may be present within their ranges. Ilexopaca var. opaca, Carpinus caroliniana, Lindera benzoin, Cornus drummondii, and <strong>Forest</strong>iera acuminata may be present in thesubcanopy. Depending on the maturity of the occurrence, physiognomy varies. The herb layer is sparse to dense.Dynamics: This community experiences frequent flooding of short duration. Very heavy siltation can kill enough individuals ofPopulus deltoides to convert the community to one dominated almost exclusively by Salix nigra. This forest often succeeds to a forestdominated by some combination of Celtis laevigata, Ulmus americana, Platanus occidentalis, Fraxinus pennsylvanica, andLiquidambar styraciflua. Regardless, succession in this community occurs relatively swiftly.Similar Associations:• Populus deltoides - Salix nigra <strong>Forest</strong> (CEGL002018) -- is more northern in distribution.Related Concepts:• Black Willow: 95 (Eyre 1980) ?• Cottonwood: 63 (Eyre 1980) ?• Eastern Cottonwood - Willow Riverfront <strong>Forest</strong> (Oberholster 1993) ?• IA7c. Eastern Cottonwood - Willow Riverfront <strong>Forest</strong> (Allard 1990) B• R1B3cI2c. Populus deltoides - Salix nigra - Celtis laevigata (Foti et al. 1994) ?• R1B3cI3a. Salix nigra (Foti et al. 1994) ?Classification Comments: This community is dominated by Populus deltoides and Salix nigra in varying proportions butcontributing at least 80% of the cover when combined. Other riverfront communities that have one or both of these species presentwill have less contribution of cover by these two species combined. Compare to its northern equivalent, Populus deltoides - Salixnigra <strong>Forest</strong> (CEGL002018). Kentucky examples may exhibit features of either type.CONSERVATION RANKING & RARE SPECIESGRank: G4G5 (1999-9-2): No informationHigh-ranked species: No informationVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>86
ELEMENT DISTRIBUTIONRange: This forest occurs in the southeastern Coastal Plain of the United States.Subnations: AL, AR, FL, GA, LA, MS, TN, TXTNC Ecoregions: 40:P, 41:C, 42:C, 43:C, 53:CUSFS Ecoregions: 231A:CC, 232F:CC, 234A:CCFederal Lands: USFS (Angelina?, Davy Crockett?, De Soto, Delta, Kisatchie, Sabine NF?, Sam Houston?, <strong>Sumter</strong>); USFWS(Chickasaw NWR, Hatchie, Lower Hatchie?, Reelfoot?)ELEMENT SOURCESReferences: Allard 1990, Ambrose 1990a, Burns and Honkala 1990b, Clark and Benforado 1981, Dickson and Segelquist 1978,Eyre 1980, Faulkner and Patrick n.d., Foti et al. 1994, Klimas 1988b, Klimas et al. 1981, McWilliams and Rosson 1990, Oberholster1993, Putnam 1951, Putnam et al. 1960, Smith 1996a, Smith and Linnartz 1980, Southeastern Ecology Working Group n.d., VanAuken and Bush 1988, Wharton 1978, Wharton et al. 1982, Wieland 1994b, Wieland 2000bRIVER BIRCH - SYCAMORE / SMOOTH ALDER / FALSE NETTLE FORESTELEMENT IDENTIFIERSNVC association: Betula nigra - Platanus occidentalis / Alnus serrulata / Boehmeria cylindrica <strong>Forest</strong>Database Code: CEGL007312Formation: Temporarily flooded cold-deciduous forest (I.B.2.N.d)Alliance: Betula nigra - (Platanus occidentalis) Temporarily Flooded <strong>Forest</strong> Alliance (A.280)ELEMENT CONCEPTSummary: This riverfront forest, dominated by Betula nigra and Platanus occidentalis, occurs primarily on levees along small riversand streams. It also is found along flowages of larger rivers ('artificial oxbows'). It ranges from Virginia to southern Georgia in thePiedmont and Coastal Plain (and into the southern and lower-elevation parts of the Southern Blue Ridge, excluding the highestelevations of the Mountains) and west to eastern Texas, and possibly the Cumberland Plateau of northern Alabama. No effort hasbeen made to subdivide this type into northern and southern, or Coastal Plain and Interior variants, although there are undoubtedlysome floristic differences between these extremes, at least in the lower strata.Platanus occidentalis may be codominant, or at least prominent, with large individuals overtopping the Betula, which tends to have agreater number of stems, but Platanus occidentalis may be more conspicuous because of its larger size. Other canopy associatesinclude Liriodendron tulipifera, Liquidambar styraciflua, Acer rubrum, Acer negundo, Ulmus americana, Ulmus rubra, Celtis spp.,and Quercus spp. The subcanopy or tall-shrub strata may include Cornus florida and Carpinus caroliniana, along with Acer rubrum,Ilex opaca, Ulmus alata, Prunus serotina, and Carya spp. Shrubs and woody vines may include Alnus serrulata, Euonymusamericana, Parthenocissus quinquefolia, Smilax rotundifolia, Toxicodendron radicans, and Vitis rotundifolia. Herbs may includeBoehmeria cylindrica, Polygonum virginianum, Rudbeckia laciniata, Sanicula sp., Symphyotrichum lateriflorum (= Asterlateriflorus), Thalictrum dioicum, Viola sororia, Polystichum acrostichoides, Woodwardia areolata, Botrychium dissectum,Botrychium virginianum, and Impatiens capensis. The exotics Ligustrum sinense, Lonicera japonica, Microstegium vimineum, andRosa multiflora may spread into disturbed examples of this community.Environment: Betula nigra requires soils near field capacity throughout the year but is relatively intolerant of flooding; Platanusoccidentalis also is intolerant of flooding during the growing season and will die if the entire tree is inundated for more than twoweeks. The absence of this community in the lower Mississippi River Alluvial Plain is attributed this intolerance, as is its mostcommon location on levees of smaller rivers. The community usually is found on the natural levee of the watercourse and, therefore,is slightly elevated from the flats behind the levee. The community is more common along small streams and blackwater streams thanalong alluvial floodplains, mostly because of the higher sustained flow rates of these larger rivers. It also is found along flowages oflarger rivers ("artificial oxbows") in southeast Kentucky. In the Upper East Gulf Coastal Plain (Shiloh <strong>National</strong> Military Park, HardinCounty, Tennessee), this association occurs along small stream floodplains.Vegetation: Betula nigra contributes at least 50%, and often more, of the tree density of stands of this community. Platanusoccidentalis may be codominant, or at least prominent, with large individuals overtopping the Betula, which tends to have a greaternumber of stems, but Platanus occidentalis may be more conspicuous because of its larger size. Other canopy associates includeLiriodendron tulipifera, Liquidambar styraciflua, Acer rubrum, Acer negundo, Ulmus americana, Ulmus rubra, Celtis spp., andQuercus spp. Pinus echinata, Pinus strobus, or Pinus virginiana may be found in some montane or submontane examples. Thesubcanopy or tall-shrub strata may include Cornus florida and Carpinus caroliniana, along with Acer rubrum, Ilex opaca, Ulmusalata, Prunus serotina, and Carya spp. Shrubs and woody vines may include Alnus serrulata, Euonymus americana, Parthenocissusquinquefolia, Smilax rotundifolia, Toxicodendron radicans, and Vitis rotundifolia. Herbs may include Boehmeria cylindrica,Polygonum virginianum, Rudbeckia laciniata, Sanicula sp., Symphyotrichum lateriflorum (= Aster lateriflorus), Thalictrum dioicum,Viola sororia, Polystichum acrostichoides, Woodwardia areolata, Botrychium dissectum, Botrychium virginianum, and Impatienscapensis. The exotics Ligustrum sinense, Lonicera japonica, Microstegium vimineum, and Rosa multiflora may spread into disturbedVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>87
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1 NatureServe is an international o
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types. However, when necessary, mod
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[Association name = floristic nomin
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Generally 5 or fewer occurrences an
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All scientific names for vascular s
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Summary: A short description of the
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Loucks, O. 1996. 100 years after Co
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ALLIANCES BY US NATIONAL VEGETATION
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virginiana, and Acer pensylvanicum
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virginiana, Juniperus virginiana va
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• Appalachian pine-oak forest (Ev
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• Maritime Oak - Holly Forest / W
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ALLIANCE CONCEPTSummary: This allia
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Related Concepts:• Yellow-Poplar:
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virginiana, Sanicula canadensis, De
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Federal Lands: COE (Claiborne Lake)
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ALLIANCE SOURCESReferences: Allard
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Quercus alba and Quercus velutina a
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USFS Ecoregions: 221Hc:CCC, 221He:C
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ALLIANCE CONCEPTSummary: This allia
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USFS Ecoregions: 221Eb:PP?, M221Ab:
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Federal Lands: NPS (Great Smoky Mou
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the associated species. More Appala
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ALLIANCE DISTRIBUTIONRange: This al
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- Page 119 and 120: USFS Ecoregions: 221Ba:CCC, 221Ha:C
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- Page 125 and 126: BIBLIOGRAPHYALNHP [Alabama Natural
- Page 127 and 128: DeYoung, H. R. 1979. The white pine
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- Page 131 and 132: Lea, C. 2002a. Vegetation classific
- Page 133 and 134: Pittman, Dr. Albert. Personal commu
- Page 135 and 136: Tobe, J. D., J. E. Fairey, III, and