community are highly disturbed and contain exotic species such as Ligustrum japonicum, Dioscorea oppositifolia, and Lonicerajaponica. Pinus echinata, in many occurrences, has been attacked by the Southern Pine Bark Beetle, which will eventually kill thetrees. The concepts of the former associations Pinus echinata - Quercus falcata / Vaccinium pallidum <strong>Forest</strong> (CEGL007494) andPinus echinata - Quercus prinus / Oxydendrum arboreum / Vaccinium pallidum <strong>Forest</strong> (CEGL007495) were merged into thisassociation and should be considered variants of this community. Pinus echinata - Quercus alba / Vaccinium pallidum / Hexastylisarifolia - Chimaphila maculata <strong>Forest</strong> (CEGL008427) includes shortleaf pine - mesic oak forests of the non-coastal plain, non-Ozark/Ouachita portion of the Pinus echinata range, with an overall more mesophytic species composition than the associationdescribed here.CONSERVATION RANKING & RARE SPECIESGRank: G3G4 (2000-1-12): This community occurs within a restricted geographic range and is uncommon within this range.Because this community is poorly known and essentially uninventoried throughout its range, there remain questions regarding itstaxonomic distinctiveness and geographic extent. Further inventory and more detailed field information may expand the current rangeand concept of this type.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This association occurs in the southern fringes of the Southern Blue Ridge, extending into the southern Ridge and Valley andCumberland Plateau. It could possibly range into the upper Piedmont.Subnations: GA, KY, NC, SC, TN?TNC Ecoregions: 50:C, 51:C, 52:?USFS Ecoregions: 231Ag:CCC, 231Dc:CCC, M221Cd:CCC, M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Great Smoky Mountains, Kings Mountain, Little River Canyon?); USFS (Chattahoochee, Cherokee?, DanielBoone, Nantahala, <strong>Sumter</strong>)ELEMENT SOURCESReferences: Allard 1990, Evans 1991, <strong>NatureServe</strong> Ecology - Southeastern U.S. unpubl. data, <strong>NatureServe</strong> Ecology - SoutheasternU.S. unpubl. data, Peet et al. unpubl. data 2002, Schafale 1998b, Southeastern Ecology Working Group n.d.SHORTLEAF PINE - POST OAK - BLACKJACK OAK / HILLSIDE BLUEBERRY WOODLANDELEMENT IDENTIFIERSNVC association: Pinus echinata - Quercus stellata - Quercus marilandica / Vaccinium pallidum WoodlandDatabase Code: CEGL003765Formation: Mixed needle-leaved evergreen - cold-deciduous woodland (II.C.3.N.a)Alliance: Pinus echinata - Quercus stellata - Quercus marilandica Woodland Alliance (A.680)ELEMENT CONCEPTSummary: This shortleaf pine - oak woodland is known from low elevations in the Southern Blue Ridge Escarpment region, andprobably extends into the adjacent southern Ridge and Valley, Cumberland Plateau, and Piedmont. This association is a woodlandwhere Pinus echinata, Quercus stellata, and Quercus marilandica occur in the canopy, each contributing 25-75% to the total canopycoverage. Other species make up less than 25% of the total canopy cover and may include Quercus falcata, Quercus coccinea, Caryapallida, Pinus rigida, and Pinus virginiana. Shrub strata vary in composition and density, but Vaccinium pallidum and otherericaceous species such as Vaccinium stamineum, Gaylussacia ursina, Gaylussacia baccata, and Kalmia latifolia are typical. Herbsfound in these woodlands include Baptisia tinctoria, Chamaecrista fasciculata (= Cassia fasciculata), Chamaecrista nictitans (=Cassia nictitans), Ceanothus americanus, Chrysopsis mariana, Clitoria mariana, Coreopsis tripteris, Crotalaria sagittalis,Desmodium rotundifolium, Saccharum giganteum, Euphorbia corollata, Helianthus atrorubens, Helianthus microcephalus, Pityopsisaspera var. adenolepis, Polygala verticillata, Rudbeckia hirta, Schizachyrium scoparium var. scoparium, Mimosa microphylla (=Schrankia microphylla), Silphium compositum, Sorghastrum nutans, Stylosanthes biflora, and Tephrosia virginiana.Environment: No informationVegetation: The canopy of stands of this association are dominated by some combination of Pinus echinata, Quercus stellata, andQuercus marilandica (each contributing 25-75% to the total canopy coverage). Other species may include Quercus falcata, Quercuscoccinea, Carya pallida, Pinus rigida, and Pinus virginiana. The shrub strata vary in composition and density, but Vacciniumpallidum and other ericaceous species such as Vaccinium stamineum, Gaylussacia ursina, Gaylussacia baccata, and Kalmia latifoliaare typical. Herbs found in these woodlands include Baptisia tinctoria, Chamaecrista fasciculata (= Cassia fasciculata),Chamaecrista nictitans (= Cassia nictitans), Ceanothus americanus, Chrysopsis mariana, Clitoria mariana, Coreopsis tripteris,Crotalaria sagittalis, Desmodium rotundifolium, Saccharum giganteum, Euphorbia corollata, Helianthus atrorubens, Helianthusmicrocephalus, Pityopsis aspera var. adenolepis, Polygala verticillata, Rudbeckia hirta, Schizachyrium scoparium var. scoparium,Mimosa microphylla (= Schrankia microphylla), Silphium compositum, Sorghastrum nutans, Stylosanthes biflora, and Tephrosiavirginiana.Dynamics: No informationVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>50
Similar Associations:• Pinus echinata - Quercus (prinus, falcata) / Oxydendrum arboreum / Vaccinium pallidum <strong>Forest</strong> (CEGL007493)• Pinus echinata / Schizachyrium scoparium Appalachian Woodland (CEGL003560) -- a related evergreen woodlandRelated Concepts:• IA6a. Dry Shortleaf Pine - Oak - Hickory <strong>Forest</strong> (Allard 1990) BClassification Comments: Probably very widespread at low elevations in and around the southern Appalachians. This associationoccurs on more xeric, extreme sites than a similar association, Pinus echinata - Quercus (prinus, falcata) / Oxydendrum arboreum /Vaccinium pallidum <strong>Forest</strong> (CEGL007493).CONSERVATION RANKING & RARE SPECIESGRank: G4? (2002-10-22): This association is probably very widespread at low elevations in and around the southernAppalachians. The final resolution of the rank and rarity of this association depends on the determination of its total global range. It isvulnerable to logging, development, invasion by exotic plant species, and fire exclusion. Shortleaf pine (Pinus echinata) populationsseem to have undergone rangewide declines in the vigor and extent. Stands of this association are threatened by the effects ofcontinued fire suppression, which would inhibit the reproduction of Pinus echinata and cause the grass-dominated herbaceous layer todeteriorate.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This shortleaf pine - oak woodland is known from low elevations in the Southern Blue Ridge Escarpment region, andprobably extends into the adjacent southern Ridge and Valley, Cumberland Plateau, and Piedmont.Subnations: GA, KY?, NC, TNTNC Ecoregions: 50:?, 51:C, 52:PUSFS Ecoregions: 221:?, 231Ag:PPP, M221Dc:CCP, M221Dd:CCCFederal Lands: NPS (Little River Canyon?); USFS (<strong>Sumter</strong>)ELEMENT SOURCESReferences: Allard 1990, Schafale and Weakley 1990, Southeastern Ecology Working Group n.d.SHORTLEAF PINE - ROCK CHESTNUT OAK / FOOTHILLS RHODODENDRON / HILLSIDE BLUEBERRY FORESTELEMENT IDENTIFIERSNVC association: Pinus echinata - Quercus prinus / Rhododendron minus / Vaccinium pallidum <strong>Forest</strong>Database Code: CEGL007496Formation: Mixed needle-leaved evergreen - cold-deciduous forest (I.C.3.N.a)Alliance: Pinus echinata - Quercus (coccinea, prinus) <strong>Forest</strong> Alliance (A.395)ELEMENT CONCEPTSummary: This association is a mixed forest occurring in association with Tallulah Falls quartzite in the Blue Ridge/Piedmonttransition of South Carolina and Georgia, at 490-610 m (1600-2000 feet) elevation. This is an open forest with a stunted canopy andscattered rock outcroppings. Pinus echinata and Quercus prinus dominate the canopy, but other species may occur, including Pinusrigida, Pinus virginiana, Quercus coccinea, Quercus falcata, Quercus stellata, and Quercus marilandica. Rhododendron minusoccurs as a tall shrub (greater than 2 m) and can be quite dense. Vaccinium pallidum is the dominant short shrub (less than 2 m), inareas where the Rhododendron is sparse or absent. Other shrubs may include Vaccinium arboreum, Rhododendron maximum, andKalmia latifolia. Smilax glauca and Vitis rotundifolia are common vines. The herbaceous stratum is sparse, with scattered grasses andforbs, including Chimaphila maculata, Pteridium aquilinum var. latiusculum, Dichanthelium dichotomum, Danthonia sericea, andSchizachyrium scoparium. This forest occurs on convex landforms over rocky, shallow soil and on broad, low ridges and exposed,convex slopes with southerly exposures, 490-610 m (1600-2000 feet) elevation.Environment: No informationVegetation: Stands are dominated by Pinus echinata and Quercus prinus, but other species may occur, including Pinus rigida, Pinusvirginiana, Quercus coccinea, Quercus falcata, Quercus stellata, and Quercus marilandica. Rhododendron minus occurs as a tallshrub (greater than 2 m) and can be quite dense. Vaccinium pallidum is the dominant short shrub (less than 2 m), in areas where theRhododendron is sparse or absent. Other shrubs may include Vaccinium arboreum, Rhododendron maximum, and Kalmia latifolia.Smilax glauca and Vitis rotundifolia are common vines. The herbaceous stratum is sparse, with scattered grasses and forbs, includingChimaphila maculata, Pteridium aquilinum var. latiusculum, Dichanthelium dichotomum, Danthonia sericea, and Schizachyriumscoparium. In an example on the Chattahoochee <strong>National</strong> <strong>Forest</strong>, the open canopy is dominated by Pinus echinata, and the midstory isdense and dominated by Quercus marilandica and Quercus prinus. The dense shrub layer is dominated by Vaccinium pallidum andalso includes Rhododendron minus, Asimina reticulata, and Kalmia latifolia. The herbaceous layer is generally sparse, but grassyopenings are present and dominated by Schizachyrium scoparium and Danthonia sericea.Dynamics: No informationSimilar Associations:Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 200451Copyright © 2004 <strong>NatureServe</strong>
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1 NatureServe is an international o
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types. However, when necessary, mod
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[Association name = floristic nomin
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Generally 5 or fewer occurrences an
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All scientific names for vascular s
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Summary: A short description of the
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- Page 45 and 46: USFS Ecoregions: 221Hc:CCC, 221He:C
- Page 47 and 48: ALLIANCE CONCEPTSummary: This allia
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- Page 65 and 66: does not occur on wet soils. It occ
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- Page 69 and 70: ELEMENT SOURCESReferences: Allard 1
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- Page 109 and 110: ELEMENT DISTRIBUTIONRange: This for
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Classification Comments: This veget
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BIBLIOGRAPHYALNHP [Alabama Natural
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DeYoung, H. R. 1979. The white pine
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Gettman, R. W. 1974. A floristic su
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Lea, C. 2002a. Vegetation classific
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Pittman, Dr. Albert. Personal commu
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Tobe, J. D., J. E. Fairey, III, and