Federal Lands: NPS (Blue Ridge Parkway?, Great Smoky Mountains); USFS (Chattahoochee, Cherokee, Daniel Boone, Nantahala,Pisgah, <strong>Sumter</strong>)ELEMENT SOURCESReferences: Allard 1990, CAP pers. comm. 1998, Eyre 1980, Peet et al. unpubl. data 2002, Schafale 1998b, Schafale and Weakley1990, Southeastern Ecology Working Group n.d.GREAT RHODODENDRON UPLAND SHRUBLANDELEMENT IDENTIFIERSNVC association: Rhododendron maximum Upland ShrublandDatabase Code: CEGL003819Formation: Hemi-sclerophyllous temperate broad-leaved evergreen shrubland (III.A.2.N.b)Alliance: Rhododendron maximum Shrubland Alliance (A.745)ELEMENT CONCEPTSummary: This community occurs along streams and on protected slopes in the mountains of North Carolina, Tennessee, SouthCarolina, Georgia, West Virginia, and Kentucky. It is a broad-leaved, evergreen shrubland which forms a continuous, dense shrubcanopy up to 5 m tall. Kalmia latifolia, Rhododendron minus, and Rhododendron catawbiense may also occur as components of theshrub stratum. Shrub vegetation beneath the upper shrub canopy may be open to dense depending on the stand's age and topographicsetting. The ground layer is dominated by leaf litter or bare soil, although scattered herbs and woody seedlings do occur. Seedlingsand saplings of Rhododendron maximum, Acer rubrum, Betula lenta, Betula alleghaniensis, and Tsuga canadensis are common andtypical herbs include Dryopteris intermedia, Heuchera villosa, Viola spp., Thelypteris noveboracensis, Listera smallii, and Galaxurceolata. This shrubland is typical along streams and on mesic, unexposed, often north-facing slopes at elevations of approximately300-1100 m (1000-3000 feet). Soils supporting this community are typically acid. Occurrences at edges of streams may flood duringrainy seasons. This community can occur as the result of disturbance and will succeed to forest with an ericaceous understory withoutsome form of disturbance. This community may have scattered woody species that are greater than 5 m tall but with generally lessthan 10% total cover.Environment: This community occurs along streams and on mesic, unexposed, often north-facing slopes at elevations ofapproximately 300-1100 m (1000-3000 feet). Soils supporting this community are typically acid. Occurrences at edges of streamsmay flood during rainy seasons.Vegetation: This evergreen, sclerophyllous shrubland is dominated by Rhododendron maximum which forms a continuous, denseshrub canopy up to 5 m tall. Kalmia latifolia, Rhododendron minus, and Rhododendron catawbiense may also occur as components ofthe shrub stratum. Shrub vegetation beneath the upper shrub canopy may be open to dense depending on the stand's age andtopographic setting. Species such as Tsuga canadensis, Pinus strobus, Acer rubrum, Betula lenta, and Liriodendron tulipifera in thetree canopy stratum make up less than 10% cover. The ground layer is dominated by leaf litter or bare soil although scattered herbsand woody seedlings do occur. Seedlings and saplings of Rhododendron maximum, Acer rubrum, Betula lenta, Betula alleghaniensis,and Tsuga canadensis are common, and typical herbs include Dryopteris intermedia, Heuchera villosa, Viola spp., Thelypterisnoveboracensis, Listera smallii, and Galax urceolata.Dynamics: Rhododendron maximum sprouts vigorously after disturbance, and this community often results from logging, fire,chestnut blight, or cessation of grazing. Stems greater than 4 cm in diameter survive hot fires, and fire generally stimulates basalsprouting, although intense annual fires may suppress reestablishment (Core 1966). Drastic overstory removal, heavy shading, anddisease have been found to decrease the density of or kill Rhododendron (Hodgdon and Pike 1961).This shrubland will become established by invading disturbed or cleared lands if there is adequate moisture and lack of directsunlight. This community can also result from secondary succession when a forest's canopy is removed (by logging, disease, etc.) andthe Rhododendron understory closes, forming a dense shrubland. The reestablishment of woody competitors is inhibited by the shadeof the dense shrub canopy as well as by phytotoxins in the litter and soil (Gant 1978). Rhododendron maximum Shrubland may persistfor over 60 years on a site (Ploucher and Carvell 1987) but will succeed to a forested community as trees that become established inthicket openings mature.Similar Associations:Related Concepts:• IC4b. Montane Rhododendron Thicket (Allard 1990) ?• Low Elevation Heath Bluff (Montane Rhododendron Subtype) (Schafale 1998b) ?• Submesotrophic Scrub (Rawinski 1992) ?Classification Comments: Rhododendron maximum Shrubland frequently occurs adjacent to wet herbaceous cliff vegetation,riparian shrublands, or within forests dominated by Tsuga canadensis, Quercus rubra, Liriodendron tulipifera, Pinus strobus,Quercus prinus, Picea rubens, or Abies fraseri. Similar ericaceous shrublands occur at higher elevations, over 1100 meters (3500feet), in the southern Appalachian Mountains. These high-elevation "heath balds" are distinguished from Rhododendron maximumUpland Shrubland by the dominance of Rhododendron catawbiense or by the occurrence of ericaceous shrubs typical of high-Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>36
elevation environments such as Leiophyllum buxifolium, Menziesia pilosa, and Photinia melanocarpa (= Aronia melanocarpa ).Disjunct populations of Rhododendron maximum are found in Maine and New Hampshire, but these populations may represent adifferent community (Hodgdon and Pike 1961).CONSERVATION RANKING & RARE SPECIESGRank: G3?Q (1998-12-14): This association is of uncertain validity and, even if valid, is of uncertain circumscription.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This community occurs in the Southern Blue Ridge, but may be possible throughout the range of Rhododendron maximum.Subnations: GA, NC, SC, TN, VA?, WVTNC Ecoregions: 49:P, 51:CUSFS Ecoregions: 221E:PP, M221A:CC, M221B:CC, M221C:CC, M221Dc:CCC, M221Dd:CCCFederal Lands: USFS (Chattahoochee?, Cherokee, Nantahala, Pisgah, <strong>Sumter</strong>)ELEMENT SOURCESReferences: Allard 1990, Core 1966, Gant 1978, Hodgdon and Pike 1961, McGee and Smith 1967, Monk et al. 1985, Nelson 1986,Phillips and Murdy 1985, Plocher and Carvell 1987, Rawinski 1992, Schafale 1998b, Southeastern Ecology Working Group n.d.TULIPTREE - SWEET BIRCH - EASTERN HEMLOCK / GREAT RHODODENDRON FORESTELEMENT IDENTIFIERSNVC association: Liriodendron tulipifera - Betula lenta - Tsuga canadensis / Rhododendron maximum <strong>Forest</strong>Database Code: CEGL007543Formation: Mixed needle-leaved evergreen - cold-deciduous forest (I.C.3.N.a)Alliance: Tsuga canadensis - Liriodendron tulipifera <strong>Forest</strong> Alliance (A.413)ELEMENT CONCEPTSummary: This association includes hemlock-hardwood forests of lower to intermediate elevations in the Southern Blue Ridge andupper Piedmont, ranging from southwestern Virginia, south and west to northwestern Georgia. These communities occur at low tomiddle elevations (1300-3500 feet) in the mountains and foothills, generally in coves, gorges, or sheltered slopes, over acid soils. Thecanopy is usually dominated by Liriodendron tulipifera or Betula lenta, but substantial portions of this community may be comprisedmainly of Tsuga canadensis and the occasional Acer rubrum. Other deciduous species more typical of "rich" coves may occur asscattered individuals; Tilia americana var. heterophylla, Fraxinus americana, and Fagus grandifolia. Other canopy/subcanopyspecies often include Quercus alba, Quercus rubra, Magnolia fraseri, Ilex opaca var. opaca, Calycanthus floridus, Halesia tetrapteravar. tetraptera, and Pinus strobus. Rhododendron maximum is scattered to dominant in the shrub stratum. Other typical shrubsinclude Kalmia latifolia and Leucothoe fontanesiana. Herbaceous cover is sparse but can be diverse and is composed of acid-lovingspecies. Typical herbs include Polystichum acrostichoides, Dennstaedtia punctilobula, Goodyera pubescens, Mitchella repens,Thelypteris noveboracensis, Galax urceolata, Viola rotundifolia, Hexastylis sp., and Tiarella cordifolia.Environment: Over its full geographic range, this association is typically found at lower to intermediate elevations (400-1060 m or1300-3500 feet) in the southern Appalachians and adjacent foothills. Habitats are located on gentle to steep, lower slopes and in covesor gorges with acidic soils. In situations where mid-slopes are in protected north-facing positions, this community can range very highup straight or even convex slopes. The type often occurs in linear patches along stream bottoms and in steep ravines in complexeswith rich cove communities. Although frequently associated with streams, it is not a wetland. Habitats in the Virginia part of the rangeare similar and are mostly situated below 900 m (3000 feet) elevation. Soils collected from plots are extremely acidic (mean pH = 3.9)and infertile, with high iron and aluminum levels and very low total base saturation.Vegetation: This association encompasses hemlock - hardwood forests with canopies dominated by mixtures of Tsuga canadensis,Liriodendron tulipifera, Betula lenta, and Acer rubrum. Other deciduous species more typical of fertile coves, including Tiliaamericana var. heterophylla, Fraxinus americana, and Fagus grandifolia, may occur as scattered individuals. Minor overstory andunderstory species include Quercus alba, Quercus rubra, Magnolia fraseri, Ilex opaca, Calycanthus floridus, Halesia tetraptera, andPinus strobus. Rhododendron maximum is scattered to dominant in the shrub stratum. Other typical shrubs include Kalmia latifoliaand Leucothoe fontanesiana. Herbaceous cover is sparse but can be diverse and is composed of acid-loving species. Typical herbsinclude Polystichum acrostichoides, Goodyera pubescens, Mitchella repens, Thelypteris noveboracensis, Galax urceolata, Hexastylisspp., and Tiarella cordifolia.Virginia examples of this association are similar to those further south but generally lack Ilex opaca, Calycanthus floridus, Halesiatetraptera, and Leucothoe fontanesiana. Presumably because of past logging, Tsuga canadensis is absent or confined to theunderstory in some stands, which have mixed canopies of Liriodendron tulipifera, Betula lenta, Acer rubrum, Magnolia acuminata,Quercus rubra, and/or Nyssa sylvatica. Hamamelis virginiana and Acer pensylvanicum are additional, frequent understory species.The shrub layers of Virginia occurrences are consistently dominated by dense (usually >50% cover), often nearly impenetrablecolonies of Rhododendron maximum. Frequent low-cover species of sparse herb layers include Galax urceolata, ChimaphilaVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>37
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1 NatureServe is an international o
- Page 11 and 12: types. However, when necessary, mod
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- Page 33 and 34: ALLIANCE CONCEPTSummary: This allia
- Page 35 and 36: Related Concepts:• Yellow-Poplar:
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- Page 39 and 40: Federal Lands: COE (Claiborne Lake)
- Page 41 and 42: ALLIANCE SOURCESReferences: Allard
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- Page 45 and 46: USFS Ecoregions: 221Hc:CCC, 221He:C
- Page 47 and 48: ALLIANCE CONCEPTSummary: This allia
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- Page 51 and 52: Federal Lands: NPS (Great Smoky Mou
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- Page 55 and 56: ALLIANCE DISTRIBUTIONRange: This al
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- Page 61 and 62: CULTIVATED FORESTEASTERN WHITE PINE
- Page 63 and 64: Federal Lands: DOD (Arnold, Fort Be
- Page 65 and 66: does not occur on wet soils. It occ
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- Page 69 and 70: ELEMENT SOURCESReferences: Allard 1
- Page 71 and 72: ELEMENT SOURCESReferences: Allard 1
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- Page 83 and 84: CONSERVATION RANKING & RARE SPECIES
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- Page 95 and 96: ELEMENT DISTRIBUTIONRange: This com
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- Page 101 and 102: ELEMENT SOURCESReferences: Allard 1
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ELEMENT DISTRIBUTIONRange: This for
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COMMON WATER-WILLOW HERBACEOUS VEGE
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• Betula nigra - Platanus occiden
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Formation: Temporarily flooded temp
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epeated flooding, this community ma
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USFS Ecoregions: 221Ba:CCC, 221Ha:C
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Classification Comments: On the Ban
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Classification Comments: This veget
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BIBLIOGRAPHYALNHP [Alabama Natural
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DeYoung, H. R. 1979. The white pine
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Gettman, R. W. 1974. A floristic su
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Lea, C. 2002a. Vegetation classific
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Pittman, Dr. Albert. Personal commu
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Tobe, J. D., J. E. Fairey, III, and