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Principles of terrestrial ecosystem ecology.pdf

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384 Glossary<br />

or from the <strong>ecosystem</strong> to groundwater or<br />

aquatic <strong>ecosystem</strong>s.<br />

Leaf area index. Leaf area per unit ground<br />

area. Projected LAI is the leaf area projected<br />

onto a horizontal plane.Total LAI is the total<br />

surface area <strong>of</strong> leaves, including the upper<br />

and lower surface <strong>of</strong> flat leaves and the<br />

cylindrical surface <strong>of</strong> conifer needles; it is<br />

approximately twice the value <strong>of</strong> projected<br />

LAI, except in the case <strong>of</strong> conifer needles,<br />

where the projected leaf area is multiplied by<br />

p (3.1416) to get total leaf area.<br />

Leaf mass ratio. Ratio <strong>of</strong> leaf mass to total<br />

plant mass.<br />

Legacy. Effect <strong>of</strong> past events on the current<br />

functioning <strong>of</strong> an <strong>ecosystem</strong>.<br />

Life history traits. Traits (e.g., seed size and<br />

number, potential growth rate, maximum<br />

size, and longevity) <strong>of</strong> an organism that<br />

determine how quickly a species can get to a<br />

site, how quickly it grows, how tall it gets, and<br />

how long it survives.<br />

Light compensation point. Light intensity at<br />

which net photosynthesis equals zero.<br />

Light-harvesting reactions. Reactions <strong>of</strong><br />

photosynthesis that transform light energy<br />

into chemical energy.<br />

Light saturation. Range <strong>of</strong> light intensities<br />

above which the rate <strong>of</strong> photosynthesis is<br />

insensitive to light intensity.<br />

Light use efficiency. Ratio <strong>of</strong> gross primary<br />

production to absorbed photosynthetically<br />

active radiation at the leaf or <strong>ecosystem</strong><br />

scale.<br />

Limitation. Reduced rate <strong>of</strong> a process (e.g., net<br />

primary production, growth or photosynthesis)<br />

due to inadequate supply <strong>of</strong> a resource<br />

(e.g., light) or low temperature.<br />

Lithosphere. Hard outermost shell <strong>of</strong> Earth.<br />

Litter. Dead plant material that is sufficiently<br />

intact to be recognizable.<br />

Litterbag. Mesh bag used to measure decomposition<br />

rate <strong>of</strong> detritus.<br />

Litterfall. Shedding <strong>of</strong> aboveground plant<br />

parts and death <strong>of</strong> plants.<br />

Littoral zone. Shore <strong>of</strong> a lake or ocean.<br />

Loam. Soil with substantial proportions <strong>of</strong> at<br />

least two size classes <strong>of</strong> soil particles.<br />

Loess. Soil derived from wind-blown silt<br />

particles.<br />

Longwave radiation. Radiation with wavelengths<br />

3000 to 30,000nm.<br />

Macr<strong>of</strong>auna. Soil animals larger than 10mm in<br />

length.<br />

Macronutrients. Nutrients that are required in<br />

large quantities by organisms.<br />

Macropores. Large pores between soil aggregates<br />

that allow rapid movement <strong>of</strong> water,<br />

roots, and soil animals.<br />

Maintenance respiration. Respiration used to<br />

support maintenance <strong>of</strong> live biomass.<br />

Mantle. Fungal hyphae that surround the<br />

root in ectomycorrhizae; also termed<br />

sheath.<br />

Mass flow. Bulk transport <strong>of</strong> solutes due to the<br />

movement <strong>of</strong> soil solution.<br />

Mass wasting. Downslope movement <strong>of</strong> soil or<br />

rock material under the influence <strong>of</strong> gravity<br />

without the direct aid <strong>of</strong> other media such as<br />

water, air, or ice.<br />

Matric potential. Component <strong>of</strong> water potential<br />

caused by adsorption <strong>of</strong> water to surfaces;<br />

it is considered a component <strong>of</strong><br />

pressure potential in some treatments.<br />

Matrix. Predominant patch type in a landscape.<br />

Mean residence time. Mass divided by the<br />

flux into or out <strong>of</strong> the pool over a given<br />

time period; synonymous with turnover<br />

time.<br />

Mechanical weathering. Physical fragmentation<br />

<strong>of</strong> the rock without chemical change.<br />

Mes<strong>of</strong>auna. Soil animals 0.2 to 10mm in<br />

length.<br />

Mesopause. Boundary between the mesosphere<br />

and thermosphere.<br />

Mesophyll cells. Photosynthetic cells in a leaf.<br />

Mesosphere. Atmospheric layer between the<br />

stratosphere and the thermosphere, which is<br />

characterized by a decrease in temperature<br />

with height.<br />

Metamorphic rocks. Sedimentary or igneous<br />

rocks that are modified by exposure to heat<br />

or pressure.<br />

Metapopulations. Populations <strong>of</strong> a species<br />

that consist <strong>of</strong> partially isolated<br />

subpopulations.<br />

Methanogen. Methane-producing bacteria.<br />

Methanotroph. Methane-consuming bacteria.

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