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Principles of terrestrial ecosystem ecology.pdf

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leached material from above, it is relatively<br />

unaffected by soil-forming processes. C horizons<br />

typically include a significant portion <strong>of</strong><br />

unweathered parent material. Finally, at some<br />

depth, there is an unweathered bedrock layer,<br />

the regolith (R).<br />

Despite the large variation among the<br />

world’s soils, they can be classed into major<br />

groups that share many <strong>of</strong> the same properties<br />

because they have formed in response to<br />

similar soil-forming factors and processes. Soil<br />

classification systems rely on the diagnostic<br />

characteristics <strong>of</strong> specific horizons and on<br />

organic matter content, base saturation, and<br />

properties that indicate wetness or dryness. The<br />

U.S. Soil Taxonomy recognizes 12 major soil<br />

groupings, called soil orders (Table 3.3) (Brady<br />

and Weil 1999). Most agronomic and <strong>ecosystem</strong><br />

studies classify soils to the level <strong>of</strong> a soil series,<br />

a group <strong>of</strong> soil pr<strong>of</strong>iles with similar pr<strong>of</strong>ile<br />

characteristics such as type, thickness, and properties<br />

<strong>of</strong> the soil horizons. Soil series can be<br />

further subdivided into types, based on the<br />

texture <strong>of</strong> the A horizon, and into phases, based<br />

on information such as landscape position,<br />

Soil Horizons and Soil Classification 59<br />

stoniness, and salinity.A comparison <strong>of</strong> soil pr<strong>of</strong>iles<br />

from the major soil orders illustrates the<br />

effect <strong>of</strong> different climatic regimes on soil<br />

development (Figs. 3.10 and 3.11).<br />

Entisols are soils with minimal soil development.<br />

They occur either because the soils are<br />

recent or processes that disrupt soil development<br />

dominate over processes that form soils.<br />

This is the most widespread soil type in the<br />

world, making up 16% <strong>of</strong> the ice-free surface.<br />

Inceptisols, in which the soil pr<strong>of</strong>ile has only<br />

begun to develop, occupy an additional 10% <strong>of</strong><br />

the ice-free surface. Rock and shifting sand<br />

account for another 14% <strong>of</strong> the ice-free surface.<br />

Thus about 40% <strong>of</strong> the ice-free surface <strong>of</strong> Earth<br />

shows minimal soil development.<br />

Histosols are highly organic soils that<br />

develop in any climate zone under conditions<br />

in which poor drainage restricts oxygen diffusion<br />

into the soil, leading to slow rates <strong>of</strong><br />

decomposition and accumulation <strong>of</strong> organic<br />

matter. There is a well-developed O horizon<br />

<strong>of</strong> undecomposed organic material where most<br />

plants are rooted.The high water table prevents<br />

the vertical leaching required for soil devel-<br />

Table 3.3. Names <strong>of</strong> the soil orders in the United States soil taxonomy and their characteristics and typical<br />

locations.<br />

Area (% <strong>of</strong><br />

Soil Order ice-free land) Major Characteristics Typical Occurrence<br />

Entisols 16.3 no well-developed horizons sand deposits, plowed fields<br />

Inceptisols 9.9 weakly developed soils young or eroded soils<br />

Histosols 1.2 highly organic; low oxygen peatland, bog<br />

Gelisols 8.6 presence <strong>of</strong> permafrost tundra, boreal forest<br />

Andisols 0.7 from volcanic ejecta; moderately developed recent volcanic areas<br />

horizons<br />

Aridisols 12.1 dry soils with little leaching arid areas<br />

Mollisols 6.9 deep dark-colored A horizon with >50% base grasslands, some deciduous forests<br />

saturation<br />

Vertisols 2.4 high content (>30%) <strong>of</strong> swelling clays; crack grassland with distinct wet and dry<br />

deeply when dry seasons<br />

Alfisols 9.7 sufficient precipitation to leach clays into a B humid forests; shrublands<br />

horizon; >50% base saturation<br />

Spodosols 2.6 sandy leached (E) horizon; acidic B horizon; cold wet climates, usually beneath<br />

surface organic accumulation conifer forests<br />

Ultisols 8.5 clay-rich B horizon, low base saturation wet tropical or subtropical climate;<br />

forest or savanna<br />

Oxisols 7.6 highly leached horizon with low clay; highly hot humid tropics beneath forests<br />

weathered on old landforms<br />

Rock and sand 14.1<br />

Data from Miller and Donahue (1990) and Brady and Weil (2001).

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