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Principles of terrestrial ecosystem ecology.pdf

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36 2. Earth’s Climate System<br />

for a past role <strong>of</strong> radiatively active gases in<br />

climate change. The unique feature <strong>of</strong> the<br />

recent anthropogenic increases in these gases<br />

is that they are occurring during an interglacial<br />

period, when Earth’s climate is already relatively<br />

warm. The Vostok record suggests that<br />

the CO2 concentration <strong>of</strong> the atmosphere is<br />

higher now than at any time in the last 400,000<br />

years. Fine-scale analysis <strong>of</strong> ice cores from<br />

Greenland suggests that large changes from<br />

glacial to interglacial climate can occur in<br />

decades or less. Such rapid transitions in<br />

the climate system to a new state may be<br />

related to sudden changes in the strength <strong>of</strong><br />

the thermohaline circulation that drives<br />

oceanic heat transport from the equator to the<br />

poles.<br />

Tree ring records, obtained from living and<br />

dead trees, provide information about the<br />

climate during the past several thousand years.<br />

The width <strong>of</strong> tree rings gives a record <strong>of</strong> temperature<br />

and moisture, and the chemical composition<br />

<strong>of</strong> wood reflects the characteristics <strong>of</strong><br />

the atmosphere at the time the wood was<br />

Time (thousands <strong>of</strong> years ago)<br />

2.7<br />

3.9<br />

8.6<br />

11.0<br />

Birch<br />

Alder<br />

Fir<br />

Pine<br />

Elm<br />

Oak<br />

0 40 0 0 0 60 0 0 40<br />

Pollen abundance (% <strong>of</strong> total tree pollen)<br />

formed. Pollen preserved in low-oxygen sediments<br />

<strong>of</strong> lakes provides a history <strong>of</strong> plant taxa<br />

and climate over the past 10,000 years or more<br />

(Fig. 2.15). Pollen records from networks<br />

<strong>of</strong> sites can be used to construct maps <strong>of</strong><br />

species distributions at various times in the<br />

past and can provide a history <strong>of</strong> species migrations<br />

across continents after climatic changes<br />

(COHMAP 1988). Other proxy records provide<br />

measures <strong>of</strong> temperature (species composition<br />

<strong>of</strong> Chironomids), precipitation (lake<br />

level), pH, and geochemistry.<br />

The combination <strong>of</strong> paleoclimate proxies<br />

indicates that climate is inherently variable<br />

over all time scales. Atmospheric, oceanic,<br />

and other environmental changes that are<br />

occurring now due to human activities must be<br />

viewed as overlays on the natural climate variability<br />

that stems from long-term changes in<br />

Earth’s surface characteristics and orbital<br />

geometry.<br />

Earth’s climate is now warmer than at<br />

any time in the last 1000 years (Fig. 2.16) and<br />

perhaps much longer.This warming is most pro-<br />

Drier,<br />

cooler<br />

climate<br />

Moister<br />

climate<br />

Drier<br />

climate<br />

Warming<br />

trend<br />

Figure 2.15. Pollen pr<strong>of</strong>ile from a bog in<br />

northwestern Minnesota showing changes<br />

in the dominant tree species over the past<br />

11,000 years. (McAndrews 1966).

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