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Catalog of the Benthic Marine Algae of the ... - SeaweedAfrica

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NUMBER 27 131<br />

Gmelin (1792:1382, 1383) erred in <strong>the</strong> same manner as<br />

Swartz, retaining <strong>the</strong> junior homonym and substituting <strong>the</strong><br />

name Fucus triangularis for <strong>the</strong> Caribbean alga (which was<br />

called Bryothamnion triangulare until Howe pointed out <strong>the</strong><br />

illegitimacy <strong>of</strong> that name). On transferring F. triqueter Linnaeus<br />

to Cystoseira, C. Agardh (1820b:61) provided a binomial<br />

(C. triquetra) that may be treated as a nomen novum<br />

in accordance with Article 72, Note 1, <strong>of</strong> <strong>the</strong> ICBN.<br />

The second pair <strong>of</strong> homonyms was Fucus articulatus S.G.<br />

Gmelin (1768:77) and F. articulatus Forsskal (1775:191).<br />

The first species was described, but not illustrated (<strong>the</strong><br />

reference to pi. II: fig. 1, apparently being erroneous,<br />

judging from <strong>the</strong> legend and a comparison <strong>of</strong> <strong>the</strong> cited<br />

figure with <strong>the</strong> description). The provenance was said to be<br />

"Oceanus Indicus." In <strong>the</strong> absence <strong>of</strong> a type specimen, a<br />

definitive identification is not possible. The Forsskal species<br />

was united with F. triqueter Linnaeus in <strong>the</strong> genus Cystoseira<br />

by C. Agardh (1820b: 61), and subsequent authors have<br />

considered <strong>the</strong> two species as congeneric, but not always<br />

conspecific. J. Agardh (1848:215-216) recognized separate<br />

species, and on transferring F. articulatus Forsskal to Cystoseira,<br />

provided a binomial (C. articulata) that may be treated<br />

as a nomen novum in accordance with Article 72, Note 1,<br />

<strong>of</strong> <strong>the</strong> ICBN. On <strong>the</strong> basis <strong>of</strong> priority or perhaps purely by<br />

chance, J.F. Gmelin (1792:1381, 1389) retained <strong>the</strong> senior<br />

homonym and substituted <strong>the</strong> name Fucus cuneiformis for<br />

F. articulatus Forsskal. By an examination <strong>of</strong> Forsskal's<br />

collection at Copenhagen (C), B0rgesen (1932:11) confirmed<br />

that Fucus articulatus Forsskal is referable to Hormophysa.<br />

Hence, Fucus cuneiformis is <strong>the</strong> earliest legitimate<br />

name for this species and its correct name is Hormophysa<br />

cuneiformis (J.F. Gmelin) P.C. Silva, new combination.<br />

Validation <strong>of</strong> <strong>the</strong> Green Algal Order Ctendocladales<br />

The order Ctenocladales was proposed by Silva<br />

(1982:147) to accommodate those green algae previously<br />

placed in <strong>the</strong> Chaetophoraceae with ulvophycean ra<strong>the</strong>r<br />

than chlorophycean ultrastructure. The nature <strong>of</strong> <strong>the</strong> publication<br />

(an encyclopedia) precluded <strong>the</strong> provision <strong>of</strong> a Latin<br />

diagnosis essential to <strong>the</strong> validity <strong>of</strong> <strong>the</strong> name. Validation is<br />

effected herewith: Chlorophytae (algae virides) filamenta<br />

uniseriata ramosa aut sarcinas cellularum constantes; thallus<br />

erectus aut prostratus aut heterotrichus, filamentis prostratis<br />

libris aut cohaerentibus; cellulae omnes nucleum unum<br />

et chloroplastum singulum parietalem laminatem plerumque<br />

pyrenoidibus thylakoidiperductis (uno aut aliquot)<br />

continentes; cellulae motiles squamatae aut nudae cum uno<br />

aut multis flagellis; flagella apicaliter affixa, symmetrica<br />

radialiter, cum systemate radicum microtubulare cruciato,<br />

radicibus rotationem contra horologii habentibus; cytokinesis<br />

per sulco cum phycoplasto nihil consociato effectus.<br />

Familia typificata: Ctenocladaceae (Borzi) Borzi<br />

(1892:48, "Ctenocladiaceae").

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