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Friesia X, 4-5

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resemble the measurements deseribed for S. avenae f. sp . tritioea,<br />

found by JOHNSON (1947) to attack barley lea ves.<br />

On barley seed the occurrence of S. nodorum has been reported<br />

by MACHACEK (1945) and NOBLE & RICHARDSON (1968) . At The Danish<br />

State Seed Testing Station pycnidia of S. nodorum were found in<br />

several seed lots in the years 1969-72 (JØRGENSEN 1973, pers. comm.) .<br />

This pap er deals with the widespread occurrence of Septoria nodo­<br />

rum and its sexual stage L eptosphaeria nodorum on barley in Den­<br />

mark, and the physiologic sp ecialization of the pathogen on ba rley<br />

and wheat .<br />

As t his apparently is the first time Septoria nodorum ha s been<br />

found associated with its sexual stage on barley, the name Lepto­<br />

sphaeria nodorum will be used in the following.<br />

OC CURRENCE OF L E P T O S P H A E R I A NODORUM<br />

ON BARLEY IN DENMARK<br />

During the growing-seasons 1970-71 disease symptoms, apparently<br />

new t o Denmark, were frequently observed on barley leaves. The<br />

symptoms were always associated with L eptosphaeria nodorum. In<br />

1972-73 the pathogen was isolated from the folIowing localities in<br />

Zealand and Jutland: Skalstrup, Snoldelev, Gadstrup, Ramsømagle,<br />

Allerslev (all in Zealand) , Jelling, Hornborg, Tvingstrup, Horsens,<br />

Langå and Randers (Jutland). At all of these localities the disease<br />

symptoms were widely occurring. The most serious attacks were<br />

found at Hornborg and Langå where 30-40 % of the barley plants<br />

were infected with a considerable loss of active leaf tissue as a conse­<br />

quence.<br />

SYMPTOMS ON BARLEY<br />

The first disease symptoms appear in Mayas brown or greyish<br />

spots on the leaf lamina and leaf sheaths or at the junction between<br />

the lamina and sheath. Later in the season lesions appear as 1-2 cm<br />

long, oval or lens-shaped, redbrown spots often continuing in chloro­<br />

tic tissue along the leafribs (Fig. 1) . More irregular or triangular<br />

formed lesions typically develop at the junction between the leaf<br />

lamina and sheath thereby affecting the entire leaf (Fig. 2) . Infected

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