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The Amphibians and Reptiles of Sinaloa, Mexico - Smithsonian ...

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152 UNIVERSITY OF KANSAS PUBLS., MUS. NAT. HIST.<br />

Specimens from western <strong>Mexico</strong> demonstrate wide overlap in<br />

characters <strong>of</strong> horridum <strong>and</strong> exasperatum. <strong>The</strong> supranasal contacts<br />

the postrostral in two <strong>of</strong> nine specimen <strong>of</strong> horridum; four <strong>of</strong> eight<br />

specimens <strong>of</strong> horridum have eight or more scales between superciliaries;<br />

the second supralabial contacts the nasal or prenasal in<br />

six <strong>of</strong> nine specimens <strong>of</strong> horridum. Although we have not seen an<br />

animal with a color pattern obviously intermediate between the<br />

two races in <strong>Sinaloa</strong>, Bogert <strong>and</strong> Martin del Campo (1956:65) suggested<br />

clinal variation in this character as well. A detailed study<br />

<strong>of</strong> Heloderma in northern <strong>Sinaloa</strong> probably will indicate that<br />

exasperatum is representative <strong>of</strong> the northern terminus <strong>of</strong> Heloderma.<br />

horridum <strong>and</strong> that clinal variation accounts for the distmctiveness<br />

<strong>of</strong> this population, as visualized by Bogert <strong>and</strong> Martin del Campo<br />

(1956:28). However, we continue to recognize two races in <strong>Sinaloa</strong><br />

until additional material from the foothills between the Rio Culiacan<br />

<strong>and</strong> the Rio Fuerte becomes available.<br />

Distribution m <strong>Sinaloa</strong>.—Known only from the area drained by<br />

the Rio Fuerte. See Fig. 57.<br />

Specimen examined.—13 km. NNE Vaca, 400 m. (* KU 80743).<br />

Literature record.—San Bias, 100 m. {Bogert <strong>and</strong> Martin del Campo,<br />

1956:32).<br />

Heloderma horridum horridum (Wiegmann)<br />

Trachyderma horridum Wiegmann, Isis von Olcen, 22:421, 1829 (type<br />

locality, "<strong>Mexico</strong>"; type locality restricted to Cuemavaca, Morelos by<br />

Bogert <strong>and</strong> Martin del Campo, Bull. Amer, Mus. Nat Hist., 109:21,<br />

April 16, 1956).<br />

Heloderma horridum Wiegmann, Isis von Oken, 22:628, 1829.<br />

Heloderma horridum horridum: Bogert <strong>and</strong> Martin del Campo, Bull. Amer.<br />

Mus. Nat Hist., 109:20, April 16,1956.<br />

Remarks.—Three specimens <strong>of</strong> Heloderma h. horridum (KU<br />

73770, 78905-06) have the following characteristics, respectively:<br />

snout-vent lengths 246, 310, 268; tail lengths 192, 227, 189; tail/<br />

snout-vent length ratios 0.78, 0.73, 0.72; ventrals from gular fold to<br />

anus 62, 64, 62; subcaudals 84, 78, 78; scales across head between<br />

posterior superciliaries 9, 7, 7; pairs <strong>of</strong> infralabials reaching chin<br />

shields 2, 1, 1; supranasal contacts postnasal no, yes, no; preanal<br />

scales not enlarged. Two specimens are essentially similar in pattern,<br />

having a brown ground color with yellow scales appearing<br />

just anterior to midbody; the yellow scales become more numerous<br />

from anterior to posterior, <strong>and</strong> there are five pairs <strong>of</strong> yellow b<strong>and</strong>s<br />

on the tail. <strong>The</strong> yellow spots begin on the nape <strong>of</strong> the other lizard<br />

(KU 78906).

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