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206 UNIVERSITY OF KANSAS PUBLS., MUS. NAT. HIST. 9080); 10.3 mi. S Culiacan (FAS 12425); 31.3 mi. NW Culiacan, Rio (SU 24078); 1.9 mi. W hwy. 15 on Culiaeancito road (SU 24079); El Dorado (AMNH 90753); 29.8 mi. N El Dorado (SU 24089); 29.9 mi. N El Dorado (SU 24088); 30.6 mi. N El Dorado (SU 24087); 31.3 mi. N El Dorado (SU 24085-86); 31.7 mi. N El Dorado (SU 24084); 32.0 mi. N El Dorado (SU 24082-83); 32.4 mi. N El Dorado (SU 24081); 9.6 mi. N Guamuchil (FAS 16998, 17016); 9.7 mi. N Mazatlan (UF 16537); 87.2 mi. N Mazatlan (FAS 12409); 6.4 mi. S Mazatlan (CAS 95755); 1.4 mi. WNW Mocorito, Rio (SU 24077); 5.4 mi. WNW Mocorito, Rio (SU 24076); 9.5 mi. WNW Mocorito, Rio (SU 24075); 10.2 mi. WNW Mocorito, Rio (SU 24074); 7.3 mi. E Navolato (SU 24080); 0.9 mi. N San Lorenzo, Rio (SU 24090); 7.1 mi. NNE Topolobampo (SU 74073); 7.4 mi. NNE Topolobampo (SU 70072). Trimorphodon lambda paucimaculata Taylor Ttimorphodon paucimaculatus Taylor, Univ. Kansas Sci. Bull., 24:527, February 16, 1938 (type locality, Mazatlan, Sinaloa). Trimorphodon lambda patidmaculata: Fugler and Dixon, Publ. Mus. Michigan State Univ., Biol. ser., 2:17, July 20, 1961. Remarks.—Taylor (1938:527) distinguished T. paucimaculatus from T. biscutatus by the presence in the former of preoculars separated from the frontal and the presence of dorsal spots "greatly elongated and fewer in number." Trimorphodon paucimaculatus is further distinguished from T. biscutatus by the absence of keels or ridges on the scales of males (Taylor, 1939b:360). Davis and Dixon (1957a:24) stated that hzmbdahas fewer ventrals and darker dorsal blotches than paucimaculatus, and that intergradation between them probably occurs in Sinaloa north of Mazadan. Duellman {1957b:240) suggested that more material may prove that the populations known as T. paucimaculatus, T, lambda, and T. biscutatus are subspecies. Fugler and Dixon (1961:17) concluded that paucimaculata is a subspecies of lambda and that the "zone of intergradation probably lies to the north of El Dorado, perhaps in the coastal plain between El Dorado and Alamos, Sonera." Fouquette and Rossman (1963:198-99) reported an intergrade with 240 ventrals from 30.7 miles north of Culiacan. Six females have 250-259 (253.8) ventrals and 73-81 (77.3) subcaudals. The dorsal scale rows at the neck are 21 in two, 22 in one, 23 in one, 24 in one, and 25 in one; the scale rows at midbody are 24 in three, 25 in two and 27 in one; the scale rows at the vent are 17 in two, 18 in two, 19 in one, and 20 in one. The supralabials are 8-9 in one and 9-9 in five; infralabials 12-12 in one, 13-13 in two, ?-13 in one, 14-13 in one, and 13-14 in one; preoculars 3-3 in three, 3-4 in one, and 3-4 in two (one preocular on right very small); postoculars 3-3 in five, and 3-4 in one; temporals 3 + 3/2 + 3 in one, 3 + 3/4 + 3 in one, 3 + 4/3 + 4 in two, and 3 + 4/3 + 5 in two. Supralabials 4-5/4-5 enter the eye in five, and 5-6/5-6 in one; the
AMPHIBIANS AND REPTILES OF SINALOA, MEXICO 207 preoculars touch the frontal in three and the preoculars are separated from the frontal in three; 21-24 (22,2) dorsal dark body blotches (KU 78930 excluded); 9-11 (9.8) dorsal dark tail blotches (KU 78930 excluded). One male (KU 37592) has 21-22-17 dorsal scale rows; the head is missing so no other counts are available. Characters of seven additional males are as follows: ventrals, 243-249 (246.9); subcaudals, 81-87 (84.3) (KU 83416 excluded); dorsal scale rows at neck, 21 in two, 22 in three, and 23 in two; scale rows at midbody, 23 in three, and 24 in four; scale rows at vent, 16 in one, 17 in two, 18 in three, and 19 in one; supralabials, 8-8 in one, 9-9 in five, and 9-10 in one; infralabials, 12-12 in three, and 13-13 in four; preoculars, 3-3 in seven; postoculars, 3-3 in six, and 4-3 in one; temporals, 2 + 3/3 + 3 in one, 3 + 3/3 + 4 in one, 3 + 4/3 + 4 in two, 3 + 3/3 + 3 in two, and 4 + 3/4 + 3 in one. Supralabials 4-5/4-5 enter the eye; the preoculars contact the frontal in one, the preoculars do not contact the frontal in six. Dorsal dark body blotches, 17-25 (21.7), and dorsal dark tail blotches, 11-14 (12.0) (KU 80765, 83416 excluded). A female and male from the vicinity of Badiraguato (KU 83415- 16) have more ventrals (251 and 248) than the intergrade reported by Fouquette and Rossman (1963:198-99). Two of the specimens examined from the southern half of Sinaloa (KU 73638, 80765) have 243 and 244 ventrals. If these specimens and one from 13 miles north of the Sinaloa-Sonora state line (Dixon, Sabbath, and Worthington, 1962:98) are intergrades, then the zone of intergradation would include most of Sinaloa northward from Mazatlan, as pointed out by Davis and Dixon (1957a :24). A zone of intergradation covering more than 500 kilometers is a condition untenable to most biologists. Although we refer all Sinaloan specimens to T. I. paucimaculata we recognize that clinal variation is probably a more reasonable explanation of evolutionary divergence in Trimofphodon in western Mexico. In addition we suspect that a detailed study of Trimorphodon biscutatus, lambda, lyrophanes, paucimaculata and vandenburghi will show that these nominal populations are all representative of a single species. Trimorphodon is a nocturnal species that is frequently found in or about rocky areas. Two specimens from San Ignacio were found about 3 meters above the ground in the rock foundation of a bridge. Distribution in Sinaloa.—Throughout the state below about 1200 meters. See Fig. 86.
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