Role <strong>of</strong> anthropogenic and natural disturbance in3establishment (0–5)Common mullein is an initial colonist in newly disturbed sites(Gross and Werner 1978, Pitcairn 2000). Seedling growth rateswere faster, producing more biomass within the same time periodon bare soils relative to vegetated soils (Gross 1984). Seedlingsdid not establish in small experimentally created openings, butthey did colonize larger openings such as those created by animaldigging. Only in the open plots did plants survive and produceseeds (Gross 1980).Current global distribution (0–5) 3Common mullein occurs throughout Europe to 64°N in Norway,east into Russia, and south to the Caucasus Mountains and tothe western Himalayas. It also is occurs in Asia Minor and China(Lid and Lid 1994, Gross and Werner 1978, Gubanov et al. 1995).Extent <strong>of</strong> the species U.S. range and/or occurrence <strong>of</strong>5<strong>for</strong>mal state or provincial listing (0–5)Common mullein occurs in nearly all <strong>of</strong> the United States(USDA 2002). In Canada it occurs mainly in Ontario, Quebec,the eastern provinces, and British Columbia (Gross and Werner1978). Verbascum tapsus is a noxious weed in Colorado, Hawaii,and Manitoba (Royer and Dickinson 1999, USDA 2002).Total <strong>for</strong> Ecological Amplitude and Distribution 16/25Feasibility <strong>of</strong> ControlScoreSeed banks (0–3) 3The seeds may remain viable <strong>for</strong> over 100 years (Kivilaan andBandurski 1981), and viable seeds have been found in soil samplesarchaeologically dated from A.D. 1300 (Ødum 1965, cited inGross and Werner 1978).Vegetative regeneration (0–3) 1<strong>Plants</strong> will not die if cut above the root crown. This will causeincreased growth <strong>of</strong> lateral branches, which will produce flowerslater (Gross and Werner 1978).Level <strong>of</strong> ef<strong>for</strong>t required (0–4) 3Common mullein is difficult to control because <strong>of</strong> the largenumber <strong>of</strong> seed and long lived seed bank. Hairs on the leavesprevent herbicides from penetrating the leaf surface. Mechanical,chemical and biological control methods can be used <strong>for</strong> commonmullein. Sowing sites with native grasses and <strong>for</strong>bs may decreaseseed germination and the chance <strong>of</strong> successful establishment.A weevil specific to common mullein was introduced to NorthAmerica from Europe. The larvae destroy up to 50% <strong>of</strong> theseeds (Gross and Werner 1978, Hoshovsky 1986, Pitcairn 2000,Rutledge and McLendon 1996).Total <strong>for</strong> Feasibility <strong>of</strong> Control 7/10Total score <strong>for</strong> 4 sections 52/100§Vicia cracca L.<strong>Ranking</strong> SummaryEcoregion known or expected to occur inSouth CoastalInterior BorealArctic AlpineYesYesYesPotential Max. ScoreEcological Impact 40 27Biological Characteristics and Dispersal 25 16Amplitude and Distribution 25 21Feasibility <strong>of</strong> Control 10 9Relative Maximum 73Climatic ComparisonCollected in<strong>Alaska</strong> regions?CLIMEXsimilarity?South Coastal Yes –Interior Boreal Yes –Arctic Alpine No YesVicia cracca has been collected in the south coastal [Seward,Ketchikan, and Unalaska] (UAM 2004) and interior boreal[Anchorage, Wasilla, Fairbanks, Rampart, and Minto] (AKNHP2003, Hultén 1968, UAM 2004), ecogeographic regions <strong>of</strong><strong>Alaska</strong>. The climatic similarity between Nome and areas wherethe species is documented has a moderate match (CLIMEX1999). There is a 77% similarity between Nome and cityChirka-Kem’, Russia, where the species occurs (Hultén 1968).Additionally, the range <strong>of</strong> bird vetch includes Røros, Norway andArkhangel’sk, Russia (Hultén 1968), which have 76% <strong>of</strong> climaticmatches with Nome respectively. This suggests that establishment<strong>of</strong> bird vetch in arctic and alpine regions <strong>of</strong> <strong>Alaska</strong> may bepossible.common names: bird vetch, cow vetchB-138Ecological ImpactScoreImpact on Ecosystem Processes (0–10) 7Bird vetch alters edaphic conditions due to fixation <strong>of</strong> atmosphericnitrogen (USDA 2002).Impact on Natural Community Structure (0–10) 7Vicia cracca can <strong>for</strong>m dense stands in <strong>Alaska</strong>. It can overgrowherbaceous vegetation and climb over shrubs, such as alder,willow, and spruce up to 2 m in height, <strong>for</strong>ming a new herbaceouslayer (Lapina pers. obs.).Impact on Natural Community Composition (0–10) 8Bird vetch quickly overtops herbaceous and low-woody species atboreal <strong>for</strong>est edges in <strong>Alaska</strong>. No data is present, but native plantspecies certainly suffer from its presence (M.L. Carlson pers obs.)Impact on Higher Trophic Levels (0–10) 5Bird vetch is highly palatable to grazing and browsing animals(USDA 2002). The seeds <strong>of</strong> bird vetch are toxic (CornelUniversity: PPID). Flowers are visited by native bees and mayalter pollination ecology <strong>of</strong> the surrounding area (Aarssen et al.1986, Klebesadel 1980, M.L. Carlson pers. obs.).Total <strong>for</strong> Ecological Impact 27/40Biological Characteristics and Dispersal ScoreMode <strong>of</strong> Reproduction (0–3) 2Bird vetch reproduces by seeds and also spreads vegetatively bygrowth <strong>of</strong> rhizomes (Aarssen et al. 1986, Klebesadel 1980, Nolen2002).Long-distance dispersal (0–3) 2The seeds <strong>of</strong> bird vetch are large and not easily dispersed. Thepods explosively split open when it dries. Plant can spread whentendrils and vine branches with seed pods cling to vectors, arebroken <strong>of</strong>f the plant, and carried to a new location (Densmore etal. 2001).
Spread by humans (0–3) 3Bird vetch was first planted in <strong>Alaska</strong> in 1909. Later it was plantedat the Fairbanks and Matanuska experiment stations where itwas evaluated <strong>for</strong> <strong>for</strong>age (Klebesadel 1980). It can be introducedwith topsoil (Densmore et al. 2001). Additionally, it can spreadalong roads on cars and heavy equipment (J. Conn pers. obs.,M. Shephard pers. obs.).Allelopathic (0–2) 0This species is not allelopathic (USDA 2002).Competitive Ability (0–3) 2Bird vetch has the ability to fix nitrogen (USDA 2002) andcompetes <strong>for</strong> resources with other species.Thicket-<strong>for</strong>ming/Smothering growth <strong>for</strong>m (0–2) 2Bird vetch overgrows herbaceous vegetation and climbs “kudzustyle”up and over shrubs such as alder and willow as well as smallspruce trees (Densmore et al. 2001).Germination requirements (0–3) 2Vicia cracca usually establishes in disturbed areas, includingthose with well-developed vegetation (Densmore et al. 2001).The seeds can easy germinate in wide range <strong>of</strong> conditions withoutscarification (J. Snyder unpubl. data).Other invasive species in the genus (0–3) 3Vicia benghalensis L. V. disperma DC., V. hirsuta (l.) S.F. Gray,V. lathyroides L., V. pannonica Crantz, V. sativa L., V. tetrasperma(L.) Schreber, and V. villosa Roth.Aquatic, wetland or riparian species (0–3) 0Bird vetch is a weed <strong>of</strong> roadsides and disturbed areas.Total <strong>for</strong> Biological Characteristics and Dispersal 16/25Ecological Amplitude and Distribution ScoreHighly domesticated or a weed <strong>of</strong> agriculture (0–4) 4In <strong>Alaska</strong>, Vicia cracca was introduced as a <strong>for</strong>age crop inFairbanks and Palmer (Densmore et al. 2001, Klebesadel 1980).Known level <strong>of</strong> impact in natural areas (0–6) 4Bird vetch has ability to invade natural areas. The species hasbeen observed growing in open mature deciduous <strong>for</strong>est nearFairbanks (Densmore et al. 2001), and it penetrates well beyondboreal <strong>for</strong>est edges in the Susitna Valley (I. Lapina, M.L. Carlsonpers. obs.). It is a significant component <strong>of</strong> grassland in northernOntario and Quebec (Aarssen et al. 1986).Role <strong>of</strong> anthropogenic and natural disturbance in3establishment (0–5)It establishes in disturbed grassy areas and along roadsides(Nolen 2002). From these areas <strong>of</strong> disturbance bird vetch caninvade habitats with moderate amounts <strong>of</strong> light penetration(M.L. Carlson pers. obs.).Current global distribution (0–5) 5Originally native to Europe, it now occurs in North America,South Africa, temperate Asia, and New Zealand (Hultén 1968).Extent <strong>of</strong> the species U.S. range and/or occurrence <strong>of</strong>5<strong>for</strong>mal state or provincial listing (0–5)Bird vetch now ranges from <strong>Alaska</strong> and British Columbia southand east across Canada to Newfoundland, south to Georgia andAlabama; a total <strong>of</strong> 36 states (USDA 2002). Vicia cracca listed asa noxious weed seed in <strong>Alaska</strong> (Group B) (<strong>Alaska</strong> AdministrativeCode).Total <strong>for</strong> Ecological Amplitude and Distribution 21/25Feasibility <strong>of</strong> ControlScoreSeed banks (0–3) 3The seeds do not germinate until the seed coat is sufficientlybroken down (by decay or abrasion) to admit water (Densmore etal. 2001). Most hard-seeded legumes have seed dormancy lasting5 years or more (M.L. Carlson pers. obs.). J. Snyder (unpubl. data)observed vetch seeds germinating without period <strong>of</strong> dormancy.Vegetative regeneration (0–3) 2There is strong vegetative growth from dormant buds <strong>of</strong>belowground roots (Aarssen et al. 1986).Level <strong>of</strong> ef<strong>for</strong>t required (0–4) 4This species is very difficult to eradicate once establishedTotal <strong>for</strong> Feasibility <strong>of</strong> Control 9/10Total score <strong>for</strong> 4 sections 73/100§B-139
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United StatesDepartment ofAgricultu
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IntroductionThe control of invasive
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Overview and aimsThe authors, repre
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The scoring from each system is ver
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While the relative ranks of species
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Figure 4. Ranks for Polygonum cuspi
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Biological Characteristics and Disp
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2.3. Potential to be spread by huma
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3.4. Current global distribution.A
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obs.), suggesting that establishmen
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DiscussionThe existing weed risk as
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AcknowledgementsThe U.S. Forest Ser
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Prather, T., S. Robins, L. Lake, an
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Appendices
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EcologicalimpactBiologicalcharacter
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Appendix A.2.Summary Scores Of Inva
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EcologicalImpactBiologicalCharacter
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Alliaria petiolata (Bieb.) Cavara &
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Biological Characteristics and Disp
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Ecological Amplitude and Distributi
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Germination requirements (0-3) 2See
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Capsella bursa-pastoris (L.) Medik.
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Spread by humans (0-3) 3The Siberia
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Known level of impact in natural ar
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Extent of the species U.S. range an
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Centaurea solstitialis L.Ranking Su
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Cirsium vulgare (Savi) TenRanking S
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Competitive Ability (0-3) 3Due to i
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Cytisus scoparius (L.) LinkRanking
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Germination requirements (0-3) 3Orc
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Digitalis purpurea L.Ranking Summar
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Galeopsis bifida Boenn. and G. tetr
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Heracleum mantegazzianumSommier & L
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Hesperis matronalis L.Ranking Summa
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Role of anthropogenic and natural d
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Known level of impact in natural ar
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Leucanthemum vulgare Lam.Ranking Su
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Competitive Ability (0-3) 2Dalmatia
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Ecological Amplitude and Distributi
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Lonicera tatarica L. common names:
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Known level of impact in natural ar
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Melilotus alba MedikusRanking Summa
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Melilotus officinalis (L.) Lam.Rank
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Allelopathic (0-2)UThere is no data
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Ecological Amplitude and Distributi
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- Page 180 and 181: Anderson, D. Phalaris. In J. C. Hic
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