Known level <strong>of</strong> impact in natural areas (0–6) 3Common St. Johnswort invades grasslands and open <strong>for</strong>estsin Cali<strong>for</strong>nia, Oregon, Washington, Idaho, and Montana. InIdaho, common St. Johnswort creates medium to dense standsin grasslands, replacing native vegetation. It has been establishedin cut and burned-over areas in Pinus ponderosa <strong>for</strong>ests in Idaho(Tisdale et al. 1959). This weed <strong>for</strong>ms large dense stands in moistgrasslands and open <strong>for</strong>est areas in British Columbia (Powellet al. 1994, White et al. 1993). Common St. Johnswort invadeslarge areas in <strong>for</strong>ests, riverbanks, and pastures in NortheasternAustralia (Parsons 1957)Role <strong>of</strong> anthropogenic and natural disturbance in3establishment (0–5)Original infestations are usually associated with logging, fire,mining, or other disturbance. It can establish in <strong>for</strong>ested areasexperiencing natural disturbances such as fire or animal diggingand burrowing (Clark 1953, Davey 1919). Vegetative propagationis usually stimulated when St. Johnswort plants are affected bygrazing, mowing, or fire (Tisdale et al. 1959).Current global distribution (0–5) 3Common St. Johnswort is native to Europe, and it is naturalizedin Asia, South Africa, Japan, North and South America, Australia,and New Zealand (Gubanov et al. 2003, USDA, ARS 2005).Extent <strong>of</strong> the species U.S. range and/or occurrence <strong>of</strong><strong>for</strong>mal state or provincial listing (0–5)Common St. Johnswort has been found in nearly all thecontinental United States and Hawaii. It is known from BritishColumbia, Ontario, Manitoba, and Quebec (Crompton et al.1988, USDA 2002). Hypericum per<strong>for</strong>atum is declared a noxiousweed in Cali<strong>for</strong>nia, Colorado, Montana, Nevada, Oregon,Washington, Wyoming, Manitoba, and Quebec (InvadersDatabase <strong>System</strong> 2003, USDA 2002).Total <strong>for</strong> Ecological Amplitude and Distribution 18/255Feasibility <strong>of</strong> ControlScoreSeed banks (0–3) 3In Australia, Clark (1953) found that St. Johnswort seeds mayremain viable in the soil <strong>for</strong> as long as 6 years. In Idaho, seedburied in soil retained viability after 3 years (Tisdale et al. 1959).Vegetative regeneration (0–3) 2Common St. Johnswort can sprout from buds on lateral roots(Rutledge and McLendon 1996).Level <strong>of</strong> ef<strong>for</strong>t required (0–4) 3Common St. Johnswort is difficult to control because <strong>of</strong> itsextensive root system and long-lived seeds. Tillage, hand pulling,mowing, or burning appears to be ineffective because vegetativereproduction may be stimulated by mechanical treatment(Tisdale et al. 1959). Common St. Johnswort can be controlledby herbicides, however, wax on the leaves inhibit herbicideuptake. Biological control has been relatively successful usingseveral leaf-feeding beetles. However, in Canada and at highelevations these insects do not thrive (Rutledge and McLendon1996, White et al. 1994).Total <strong>for</strong> Feasibility <strong>of</strong> Control 8/10Total score <strong>for</strong> 4 sections 52/100§B-56
Impatiens glandulifera Royle common names: ornamental jewelweed,policemen’s helmet, Himalayan balsam, Washington orchid<strong>Ranking</strong> SummaryEcoregion known or expected to occur inSouth CoastalInterior BorealArctic AlpineYesYesNoPotential Max. ScoreEcological Impact 40 29Biological Characteristics and Dispersal 23 22Amplitude and Distribution 25 22Feasibility <strong>of</strong> Control 10 7Relative Maximum 82Climatic ComparisonCollected in<strong>Alaska</strong> regions?CLIMEXsimilarity?South Coastal Yes –Interior Boreal Yes –Arctic Alpine No NoOrnamental jewelweed has been recorded in Haines (AKEPIC2004) and Wrangell (M. Shephard pers. com.). It is widely plantedas an ornamental in Anchorage (I. Lapina pers. obs.). The length<strong>of</strong> the growing season may be a limiting factor in its northerndistribution, while absolute minimum temperatures appear to benot significantly limiting. Beerling (1993) calculated a minimumrequired value <strong>of</strong> 2,195 day-degrees from its present distributionin Europe and used this to predict the northward spread. Thegrowing season in arctic alpine <strong>Alaska</strong> is less than 2,195 daydegrees:1,112 day-degrees in Nome, 1,564 in Dillingham, 313 inBarrow (WRCC 2001). This suggests that Impatiens glanduliferacannot extend its distribution into arctic alpine <strong>Alaska</strong>.Ecological ImpactScoreImpact on Ecosystem Processes (0–10) 7This plant can alter waterflow and increase erosion and flooding athigh densities (King County 2004). Additionally, as it suppressesthe growth <strong>of</strong> co-occurring species it likely reduces availableresources (light, nutrients, moisture) (Prots and Klotz 2004)Impact on Natural Community Structure (0–10) 8Impatiens gladulifera creates a dense canopy, eliminating mostlayers below. Despite being an annual, its dry stems persist asa layer the following spring (Beerling and Perrins 1993, KingCounty 2004).Impact on Natural Community Composition (0–10) 7This aggressive plant is able to reduce the growth <strong>of</strong> native species,eventually replacing them at sites where it gets established (KingCounty 2004, Prots and Klotz 2004). In studies in Great Britainvery few species were found co-occurring with ornamentaljewelweed (Beerling and Perrins 1993,).Impact on Higher Trophic Levels (0–10) 7This plant competes with native plants <strong>for</strong> pollinators reducingseed set in native plants. Pollinators include several species <strong>of</strong>bumblebees, honeybees, moths, and wasps (Beerling and Perrins1993, Chittka and Schürkens 2001, King County 2004). It altershabitats <strong>for</strong> wildlife species. Because <strong>of</strong> high holocellulose contentin its stems, it persists as a litter the following spring, suppressingcompeting seedlings <strong>of</strong> other species (Beerling and Perrins 1993).Nectar <strong>of</strong> Impatiens glandulifera is rich and more rewarding thanthat <strong>of</strong> any known native plant in Central Europe (Chittka andSchürkens 2001).Total <strong>for</strong> Ecological Impact 29/40B-57Biological Characteristics and Dispersal ScoreMode <strong>of</strong> Reproduction (0–3) 3Jewelweed reproduces entirely by seeds. Medium-sized plantsgrowing at a density <strong>of</strong> 20 per square meter produce between700 and 800 seeds (Beerling and Perrins 1993). Large plants canproduce up to 2,500 seeds, (Chittka and Schürkens 2001, KingCounty 2004).Long-distance dispersal (0–3) 3The seeds can be transported long distance by both water andsmall mammals (Beerling and Perrins 1993, King County 2004).The rate <strong>of</strong> spread in the U.K. was estimated as 2–5 km per year(Beerling and Perrins 1993).Spread by humans (0–3) 3Ornamental jewelweed is a garden plant that has escapedcultivation. It is frequently sold at nurseries (King County 2004),and commonly planted in southern <strong>Alaska</strong>.Allelopathic (0–2)UThere is no record <strong>of</strong> allelopathy.Competitive Ability (0–3) 3This species is an aggressive competitor, overtopping andsuppressing the growth <strong>of</strong> neighboring species (Beerling andPerrins 1993). Impatiens glandulifera tolerates many types <strong>of</strong> soil,it occurs on fine and coarse alluvium, maritime shingle, freedrainingmineral soils, and peat. It can grow in full sun as well aspartial shade. It has been found along industrial rivers, suggestingit is tolerant or resistant to aquatic and/or atmospheric pollution(Beerling and Perrins 1993, King County 2004).Thicket-<strong>for</strong>ming/Smothering growth <strong>for</strong>m (0–2) 2Impatiens glandulifera creates dense thickets up to 10 feet talland it is normally taller than surrounding herbaceous vegetation(Beerling and Perrins 1988).Germination requirements (0–3) 2This plant requires open soil to germinate and establish (Beerlingand Perrins 1993). It will also germinate in tidal wrack (M.Shephard pers. obs.).Other invasive species in the genus (0–3) 3Impatiens walleriana Hook. f. is considered an invasive in Hawaii(USDA 2002). Impatiens parviflora DC. is an Asiatic speciesinvasive in northern Europe (Lid and Lid 1994)Aquatic, wetland or riparian species (0–3) 3Jewelweed is found in wetlands, riparian areas, streamsides,lowlands, wet meadows and <strong>for</strong>ests, and roadside ditches. It isplanted in gardens and parks (Beerling and Perrins 1993, KingCounty 2004).Total <strong>for</strong> Biological Characteristics and Dispersal 22/23Ecological Amplitude and Distribution ScoreHighly domesticated or a weed <strong>of</strong> agriculture (0–4) 4Ornamental jewelweed is known as “one <strong>of</strong> the many desirablehardy plants” <strong>for</strong> use in flower gardens (Beerling and Perrins1993).
- Page 1:
United StatesDepartment ofAgricultu
- Page 5 and 6:
IntroductionThe control of invasive
- Page 7 and 8:
Overview and aimsThe authors, repre
- Page 9 and 10:
The scoring from each system is ver
- Page 11 and 12:
While the relative ranks of species
- Page 13 and 14:
Figure 4. Ranks for Polygonum cuspi
- Page 15 and 16:
Biological Characteristics and Disp
- Page 17 and 18:
2.3. Potential to be spread by huma
- Page 19 and 20:
3.4. Current global distribution.A
- Page 21 and 22:
obs.), suggesting that establishmen
- Page 23 and 24:
DiscussionThe existing weed risk as
- Page 25 and 26:
AcknowledgementsThe U.S. Forest Ser
- Page 27 and 28:
Prather, T., S. Robins, L. Lake, an
- Page 29:
Appendices
- Page 32 and 33:
EcologicalimpactBiologicalcharacter
- Page 34 and 35:
Appendix A.2.Summary Scores Of Inva
- Page 36 and 37:
EcologicalImpactBiologicalCharacter
- Page 38 and 39:
Alliaria petiolata (Bieb.) Cavara &
- Page 40 and 41:
Biological Characteristics and Disp
- Page 42 and 43: Ecological Amplitude and Distributi
- Page 44 and 45: Feasibility of ControlScoreSeed ban
- Page 46 and 47: Germination requirements (0-3) 2See
- Page 48 and 49: Capsella bursa-pastoris (L.) Medik.
- Page 50 and 51: Spread by humans (0-3) 3The Siberia
- Page 52 and 53: Known level of impact in natural ar
- Page 54 and 55: Extent of the species U.S. range an
- Page 56 and 57: Centaurea solstitialis L.Ranking Su
- Page 58 and 59: Feasibility of ControlScoreSeed ban
- Page 60 and 61: Cirsium vulgare (Savi) TenRanking S
- Page 62 and 63: Competitive Ability (0-3) 3Due to i
- Page 64 and 65: Feasibility of ControlScoreSeed ban
- Page 66 and 67: Cytisus scoparius (L.) LinkRanking
- Page 68 and 69: Germination requirements (0-3) 3Orc
- Page 70 and 71: Digitalis purpurea L.Ranking Summar
- Page 72 and 73: Extent of the species U.S. range an
- Page 74 and 75: Feasibility of ControlScoreSeed ban
- Page 76 and 77: Galeopsis bifida Boenn. and G. tetr
- Page 78 and 79: Extent of the species U.S. range an
- Page 80 and 81: Heracleum mantegazzianumSommier & L
- Page 82 and 83: Hesperis matronalis L.Ranking Summa
- Page 84 and 85: Role of anthropogenic and natural d
- Page 86 and 87: Feasibility of ControlScoreSeed ban
- Page 88 and 89: Biological Characteristics and Disp
- Page 90 and 91: Competitive Ability (0-3) 3Hydrilla
- Page 94 and 95: Known level of impact in natural ar
- Page 96 and 97: Role of anthropogenic and natural d
- Page 98 and 99: Feasibility of ControlScoreSeed ban
- Page 100 and 101: Leucanthemum vulgare Lam.Ranking Su
- Page 102 and 103: Competitive Ability (0-3) 2Dalmatia
- Page 104 and 105: Ecological Amplitude and Distributi
- Page 106 and 107: Lonicera tatarica L. common names:
- Page 108 and 109: Other invasive species in the genus
- Page 110 and 111: Known level of impact in natural ar
- Page 112 and 113: Biological Characteristics and Disp
- Page 114 and 115: Ecological Amplitude and Distributi
- Page 116 and 117: Melilotus alba MedikusRanking Summa
- Page 118 and 119: Melilotus officinalis (L.) Lam.Rank
- Page 120 and 121: Allelopathic (0-2)UThere is no data
- Page 122 and 123: Ecological Amplitude and Distributi
- Page 124 and 125: Biological Characteristics and Disp
- Page 126 and 127: Other invasive species in the genus
- Page 128 and 129: Role of anthropogenic and natural d
- Page 130 and 131: Plantago major L.Ranking SummaryEco
- Page 132 and 133: Competitive Ability (0-3) 1Annual b
- Page 134 and 135: Poa pratensis ssp. pratensis L.comm
- Page 136 and 137: Polygonum aviculare L. common names
- Page 138 and 139: Competitive Ability (0-3) 2Black bi
- Page 140 and 141: Other invasive species in the genus
- Page 142 and 143:
Known level of impact in natural ar
- Page 144 and 145:
Feasibility of ControlScoreSeed ban
- Page 146 and 147:
Rumex acetosella L.Ranking SummaryE
- Page 148 and 149:
Long-distance dispersal (0-3) 3The
- Page 150 and 151:
Current global distribution (0-5) 3
- Page 152 and 153:
Long-distance dispersal (0-3) 3Ragw
- Page 154 and 155:
Feasibility of ControlScoreSeed ban
- Page 156 and 157:
Sonchus arvensis L. common names: f
- Page 158 and 159:
Spread by humans (0-3) 3European mo
- Page 160 and 161:
Ecological Amplitude and Distributi
- Page 162 and 163:
Stellaria media (L.) Vill.Ranking S
- Page 164 and 165:
Taraxacum officinale ssp. officinal
- Page 166 and 167:
Aquatic, wetland or riparian specie
- Page 168 and 169:
Trifolium hybridum L.Ranking Summar
- Page 170 and 171:
Current global distribution (0-5) 3
- Page 172 and 173:
Long-distance dispersal (0-3) 2The
- Page 174 and 175:
Role of anthropogenic and natural d
- Page 176 and 177:
Vicia villosa RothRanking SummaryEc
- Page 178 and 179:
Current global distribution (0-5) 0
- Page 180 and 181:
Anderson, D. Phalaris. In J. C. Hic
- Page 182 and 183:
Best, K.F., G.G. Bowes, A.G. Thomas
- Page 184 and 185:
Cameron, E. 1935. A study of the na
- Page 186 and 187:
Corbin, J.D., M. Thomsen, J. Alexan
- Page 188 and 189:
Douglas, G.W. and A. MacKinnon. 199
- Page 190 and 191:
Frankton, C. and G.A. Mulligan. 197
- Page 192 and 193:
Haggar, R.J. 1979. Competition betw
- Page 194 and 195:
Howard, J.L. 2002. Descurainia soph
- Page 196 and 197:
Klinkhamer, P.G. and T.J. De Jong.
- Page 198 and 199:
MAFF - Ministry of Agriculture, Foo
- Page 200 and 201:
Miki, S. 1933. On the sea-grasses i
- Page 202 and 203:
Paddock, Raymond, E. III. Environme
- Page 204 and 205:
Proctor, V.W. 1968. Long-distance d
- Page 206 and 207:
Saner, M.A., D.R. Clements, M.R. Ha
- Page 208 and 209:
Stebbins, L.G. 1993. Tragopogon: Go
- Page 210 and 211:
Townshend, J.L. and T.R. Davidson.
- Page 212 and 213:
Washington State Department of Ecol
- Page 214 and 215:
Wolfe-Bellin, K.S. and K.A. Moloney
- Page 216 and 217:
B. Invasiveness Ranking1. Ecologica
- Page 218 and 219:
2.5. Competitive abilityA. Poor com
- Page 220:
4. Feasibility of Control4.1. Seed