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atoll research bulletin no. 392 the flora of - Smithsonian Institution ...

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1 to 6 cm wide, very slenderly lanceolate-elliptic, or obovate-lanceolate, acuminate,<br />

deeply separated, dark to light green and glabrous above, glaucous and pale beneath,<br />

sometimes red-tinged, sometimes slightly hairy beneath, midrib green to deep red;<br />

petioles, 5 to 32 cm long, green to deep red; stipules, usually with 3 to 5 lanceolate lobes<br />

to 1 cm long, deciduous; flowers, about 1 to 2 cm across, mo<strong>no</strong>ecious, petal-less, <strong>of</strong>ten<br />

<strong>no</strong>t developing, pendulous, borne in few-flowered, lax, axillary paniculate clusters, 3 to<br />

10 cm long, with male and female flowers in <strong>the</strong> same inflorescence, <strong>the</strong> latter near <strong>the</strong><br />

base; bracts, linear, deciduous; sepals 5, pale yellow or tinged with red, glabrous<br />

without, puberulent within; male flowers, on pedicels 0.5 to 1 cm long, calyx, 3 to 8 mm<br />

long, campanulate with 5 triangular lobes extending to middle <strong>of</strong> calyx, stamens 10 in 2<br />

whorls, alternately short and long, filaments free, an<strong>the</strong>rs small, basal disk orange, fleshy<br />

with 5 double lobes; female flowers, on pedicels 1 to 2.5 cm long, usually larger than<br />

males, calyx 5-partite to base, 1 cm long or more, ovary, 3 to 4 mm long, 3-carpellary,<br />

6-ridged, glabrous, greenish-white with pink or red streaks, on a 10-lobed disc, style<br />

connate, surmounted by 3 lobed white stigmas, each lobe much divided; fruit, 1.5 to 2<br />

cm long, subglobose to ellipsoid capsule with narrow longitudinal wings or ridges, 3-<br />

seeded, splitting explosively; seeds, about 12 mm long, ellipsoid, smooth, gray mottled<br />

with dark blotches, carunculate, with a thin crustaceous testa; tubers, developing as<br />

swellings on adventitious roots at a short distance from <strong>the</strong> stem, usually 4 to 10 per<br />

plant, 10 to 100 cm long and 3 to 15 cm in diameter, usually weighing 1 to 10 kg or<br />

more, cylindrical or tapering, occasionally branched, outer skin, rough or smooth, light<br />

to darker brown to white or pinkish, inner rind or cortex usually white, but sometimes<br />

tinged with pink or brown, core or pith consisting <strong>of</strong> rich white or yellowish starch with<br />

few xylem bundles and latex tubes. Food plant in I Kiribati and Tuvaluan gardens at<br />

Location and Topside workshops. This easy-to-grow plant, which thrives on <strong>the</strong> poorest<br />

<strong>of</strong> soils, has become <strong>the</strong> dominant staple root crop, displacing taro and yam as <strong>the</strong><br />

dominant staple in many areas <strong>of</strong> high-island Micronesia, and in Fiji, Tonga, parts <strong>of</strong> <strong>the</strong><br />

Cook Islands, and in <strong>the</strong> drier areas <strong>of</strong> Vanuatu, New Caledonia, and Papua New<br />

Guinea. 5, 6, 7.<br />

Pedilanthus tithymaloides (L.) Poit. "slipper flower", "slipper spurge", "shoe spurge",<br />

"zigzag plant", "redbird cactus", "ribbon cactus"<br />

syns. P. carinatus Spreng.; Euphorbia tithymaloides L.<br />

Recent introduction. Caribbean. Occasional. Erect or somewhat decumbent,<br />

sparsely branching, slightly succulent subshrub, up to 1 m or higher, with white latex and<br />

fleshy, cylindrical stems which zigzag from <strong>no</strong>de to <strong>no</strong>de; leaves, 5 to 10 cm long,<br />

alternate, soon falling, fleshy, waxy, ovate to lanceolate, subsessile, <strong>of</strong>ten variegated or<br />

streaked green, white, yellowish or pink tinted, or in some forms completely albi<strong>no</strong>;<br />

-flowers,me<strong>no</strong>ec~u~,~~spi~s,-p~l~e~-yellaw.is~or~n.~c<br />

terminal cymose clusters; cyathium, about 12 mm long, narrow, boat- or slipper-shaped<br />

red or pink bracts containing 1 female and several male flowers, with 4 glands; fruit, 7.5<br />

by 9 mm, capsular; seeds, 5 mm long, ovate. Planted ornamental. 3(58681), 5, 6, 7.

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