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Fishery bulletin of the Fish and Wildlife Service - NOAA

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FISHERY BULLETIN OF THE FISH AND WILDLIFE SERVICE 275<br />

migrate seaward. At <strong>the</strong> end <strong>of</strong> <strong>the</strong> migration period a part <strong>of</strong> this population is<br />

present in <strong>the</strong> lake. These fish remain for ano<strong>the</strong>r year <strong>and</strong> obtain more growth. In<br />

<strong>the</strong> fourth year, <strong>the</strong> largest individuals remaining in <strong>the</strong> lake proceed seaward, <strong>the</strong><br />

time <strong>of</strong> appearance in <strong>the</strong> migration being correlated with <strong>the</strong>ir size. The slowest<br />

growing individuals <strong>of</strong> <strong>the</strong> entire progeny which have not migrated remam in <strong>the</strong> lake<br />

for ano<strong>the</strong>r year <strong>and</strong> <strong>the</strong>n migrate seaward in <strong>the</strong>ir fifth year.<br />

The older fish are <strong>of</strong> a larger average size than those <strong>of</strong> lesser age <strong>and</strong> <strong>the</strong>ir larger<br />

size is due to <strong>the</strong> longer growing period that precedes migration. <strong>Fish</strong> in <strong>the</strong> older<br />

age groups are usually <strong>the</strong> slower growing fish <strong>of</strong> <strong>the</strong> progeny from a particular spawning.<br />

Thus, <strong>the</strong> urge to migrate seaward is related to <strong>the</strong> size <strong>and</strong> growth rate <strong>of</strong><br />

fingerlings, <strong>and</strong> it appears that environmental conditions that affect <strong>the</strong> growth <strong>of</strong> <strong>the</strong><br />

fish during <strong>the</strong> time spent in <strong>the</strong> lake also affect <strong>the</strong> tune at which <strong>the</strong> fingerlings<br />

migrate to <strong>the</strong> ocean.<br />

The data on <strong>the</strong> percentage <strong>of</strong> males in <strong>the</strong> samples <strong>of</strong> migrants which were examined<br />

to determine sex are presented in table 28. The males <strong>and</strong> females were<br />

equally represented. Grouping <strong>the</strong> 3- <strong>and</strong> 4-year fish, it was found that <strong>the</strong> total <strong>of</strong><br />

11,080 fish examined consisted <strong>of</strong> 5,557 males <strong>and</strong> 5,523 females. The slight variations<br />

in <strong>the</strong> sex ratios from year to year are probably due to chance because <strong>the</strong>re is<br />

no significant statistical difference in <strong>the</strong> ratios.<br />

TABLE 28.—Number <strong>of</strong> 8-year <strong>and</strong> 4-year migrants examined <strong>and</strong> percentage <strong>of</strong> males in <strong>the</strong> samples<br />

1925<br />

1926<br />

1927 . ..<br />

1928<br />

1929<br />

1930<br />

1931<br />

1932<br />

1933<br />

1934. . _<br />

Total<br />

Year<br />

Number <strong>of</strong><br />

3-year fish<br />

examined<br />

£70<br />

448<br />

211<br />

491<br />

318<br />

1,308<br />

1,764<br />

1,286<br />

646<br />

802<br />

7, 804<br />

Number <strong>of</strong><br />

males<br />

296<br />

232<br />

115<br />

262<br />

168<br />

659<br />

831<br />

632<br />

320<br />

401<br />

8 916<br />

Percentage<br />

<strong>of</strong> males<br />

51.9<br />

51.8<br />

54.5<br />

53.4<br />

52.8<br />

60.4<br />

47.4<br />

60.3<br />

49.5<br />

50.0<br />

60 18<br />

SEX RATIOS OF ADULT FISH<br />

Number <strong>of</strong><br />

4-year fish<br />

examined<br />

72<br />

160<br />

144<br />

224<br />

287<br />

674<br />

277<br />

833<br />

626<br />

90<br />

3 276<br />

Number <strong>of</strong><br />

males<br />

40<br />

71<br />

75<br />

127<br />

161<br />

335<br />

132<br />

401<br />

262<br />

47<br />

1,641<br />

Percentage<br />

<strong>of</strong> males<br />

The sex ratio <strong>of</strong> <strong>the</strong> adult fish is in marked contrast to that <strong>of</strong> <strong>the</strong> seaward migrants,<br />

Data on <strong>the</strong> percentage occurrence <strong>of</strong> males in <strong>the</strong> samples for <strong>the</strong> years 1922 <strong>and</strong> 1924<br />

to 1936 are presented in table 29, arranged according to <strong>the</strong> length <strong>of</strong> time spent in<br />

<strong>the</strong> ocean. The percentage occurrence <strong>of</strong> <strong>the</strong> males decreases with increased ocean<br />

residence. All <strong>of</strong> <strong>the</strong> zero-ocean fish 8 are males. The average percentages <strong>of</strong> males<br />

in <strong>the</strong> one-ocean fish, <strong>of</strong> varying .periods <strong>of</strong> fresh-water residence, range from 100<br />

percent to 75 percent. The average percentages <strong>of</strong> males in <strong>the</strong> two-ocean fish range<br />

from 02 percent to 32 percent, while <strong>the</strong> average percentages <strong>of</strong> males hi <strong>the</strong> threeocean<br />

fish <strong>of</strong> varying fresh-water residence range from 38 percent to 35 percent.<br />

• <strong>Fish</strong> which spend only a few months In <strong>the</strong> ocean <strong>and</strong> return as mature flshln <strong>the</strong> fall <strong>of</strong> <strong>the</strong> same year in which <strong>the</strong>y migrated<br />

seaward.<br />

65.6<br />

47.3<br />

62.1<br />

56.7<br />

66.1<br />

61.3<br />

47.6<br />

48.1<br />

48.0<br />

52.2<br />

60 09

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