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Fishery bulletin of the Fish and Wildlife Service - NOAA

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356 FISHERY BULLETIN OF THE FISH AND WILDLIFE SERVICE<br />

ments <strong>of</strong> <strong>the</strong>se two concentration zones are such as to label <strong>the</strong>ir existence as a real<br />

phenomenon, <strong>and</strong> not a chance result to be ascribed to inadequate data. However, <strong>the</strong><br />

mere presence <strong>of</strong> two distinct groupings <strong>of</strong> whitefish throughout all or most <strong>of</strong> <strong>the</strong><br />

season does not make absolutely necessary <strong>the</strong> assumption <strong>of</strong> two permanently separated<br />

stocks. It is possible that conditions within <strong>the</strong> lake at certain seasons may produce<br />

an "ecological division" <strong>of</strong> an o<strong>the</strong>rwise homogeneous population.<br />

Records <strong>of</strong> a number <strong>of</strong> vertical series <strong>of</strong> temperature readings made in nor<strong>the</strong>astern<br />

Lake Michigan 88 failed to give a clue to <strong>the</strong> causé <strong>of</strong> two zones <strong>of</strong> concentration <strong>of</strong><br />

whitefish. Both <strong>the</strong> inshore <strong>and</strong> <strong>of</strong>fshore concentrations <strong>of</strong> legal fish were below <strong>the</strong><br />

<strong>the</strong>rmocline 8 * in June, July, <strong>and</strong> August <strong>and</strong> hence were in a region with extremely<br />

small temperature gradients. Preferences for water <strong>of</strong> different temperature, <strong>the</strong>refore,<br />

do not provide a logical explanation for <strong>the</strong> presence <strong>of</strong> two concentrations. The<br />

illegal whitefish <strong>of</strong> <strong>the</strong> inshore concentration were in <strong>the</strong> region <strong>of</strong> <strong>the</strong> <strong>the</strong>rmocline in<br />

July <strong>and</strong> August, hence in substantially warmer water than were <strong>the</strong> fish <strong>of</strong> <strong>the</strong> <strong>of</strong>fshore<br />

concentration. However, both groups were below <strong>the</strong> <strong>the</strong>rmocline in June, <strong>and</strong> an<br />

inshore concentration at <strong>the</strong> <strong>the</strong>rmocline was lacking in September.<br />

Important arguments in support <strong>of</strong> <strong>the</strong> assumption <strong>of</strong> <strong>the</strong> existence <strong>of</strong> inshore <strong>and</strong><br />

deep-water populations <strong>of</strong> whitefish are:<br />

(1) The separation into two groups involved both large (legal) <strong>and</strong> small (illegal)<br />

fish. Consequently, <strong>the</strong> two groups are not entirely <strong>the</strong> result <strong>of</strong> different reactions <strong>of</strong><br />

fish <strong>of</strong> different size to <strong>the</strong> same or similar environmental factors. This statement<br />

holds even though <strong>the</strong> concentration zones <strong>of</strong> <strong>the</strong> legal <strong>and</strong> illegal fish were not always<br />

identical in <strong>the</strong> same month.<br />

(2) The fish <strong>of</strong> both concentration areas have similar seasonal vertical movements.<br />

The similarity <strong>of</strong> vertical movements kept <strong>the</strong> two zones <strong>of</strong> concentration distinct in all<br />

months but September. The presence <strong>of</strong> only one peak in <strong>the</strong> September data may<br />

represent <strong>the</strong> temporary approximation <strong>of</strong> <strong>the</strong> two concentrations or may be <strong>the</strong> result<br />

<strong>of</strong> lack <strong>of</strong> information on <strong>the</strong> distribution <strong>of</strong> whitefish beyond <strong>the</strong> 110-foot contour.<br />

(3) There is evidence that some whitefish seldom, if ever, spawn in shallow wa'ter.<br />

The introduction <strong>of</strong> <strong>the</strong> deep trap net on gill-net grounds or in areas beyond <strong>the</strong> reach<br />

<strong>of</strong> pound nets was marked by <strong>the</strong> capture <strong>of</strong> considerable numbers <strong>of</strong> whitefish <strong>of</strong> exceptionally<br />

large size. These large fish could not be taken on <strong>the</strong>se same grounds by<br />

<strong>the</strong> gill nets commonly employed since <strong>the</strong>ir great size prevented <strong>the</strong>ir becoming gilled.<br />

Pound nets, which are selective only with respect to small fish, are fully capable <strong>of</strong><br />

taking large individuals <strong>of</strong> any size. Consequently, <strong>the</strong>ir failure to capture many fish<br />

as large as those found in <strong>the</strong> early catches <strong>of</strong> <strong>the</strong> deep trap nets may be taken as evidence<br />

that <strong>the</strong>se giant individuals were seldom, if ever, present on <strong>the</strong> inshore pound-net<br />

grounds, at least during <strong>the</strong> period <strong>of</strong> fishing operation.<br />

It must be remembered, never<strong>the</strong>less, that <strong>the</strong>re is no pro<strong>of</strong> that <strong>the</strong> smaller mature<br />

fish <strong>of</strong> <strong>the</strong> <strong>of</strong>fshore group <strong>of</strong> whitefish do not spawn in shallow water. The separation<br />

<strong>of</strong> <strong>the</strong> whitefish into two depth groups may represent only a summer <strong>and</strong> early-autumn<br />

condition. Possibly most <strong>of</strong> <strong>the</strong> small fish <strong>of</strong> both groups spawn in shallow water <strong>and</strong><br />

most <strong>of</strong> <strong>the</strong> large fish <strong>of</strong> both groups spawn in deeper water. However, it also seems<br />

logical to hold that <strong>the</strong> giant fish taken in deep trap nets were members <strong>of</strong> a deepwater<br />

population (that lived beyond <strong>the</strong> reach <strong>of</strong> pound nets) that had survived to a<br />

size at which <strong>the</strong>y could not be taken in gill nets, <strong>and</strong> hence had become exempt from<br />

capture in <strong>the</strong> commercial fishery.<br />

Even if <strong>the</strong> inshore <strong>and</strong> <strong>of</strong>fshore groups <strong>of</strong> whitefish are held to be semi-independent<br />

or independent, it must be recognized that both groups exhibit similar fluctuations in <strong>the</strong><br />

fishery. The records <strong>of</strong> <strong>the</strong> catch per lift <strong>and</strong> <strong>of</strong> production in M-3 (table 17 <strong>and</strong> appendix<br />

B) demonstrate a close correlation between <strong>the</strong> annual fluctuations in <strong>the</strong><br />

» Temperature data were not available trom <strong>the</strong> north channel (region north <strong>of</strong> <strong>the</strong> Beaver bl<strong>and</strong> archipelago), <strong>the</strong> center <strong>of</strong> <strong>the</strong> deep-trap-net<br />

fishery. However, <strong>the</strong> relatively limited local variation in. temperature conditions at stations sou<strong>the</strong>ast, south, <strong>and</strong> northwest <strong>of</strong> Beaver bl<strong>and</strong> <strong>and</strong><br />

sou<strong>the</strong>ast <strong>of</strong> Manietique suggests that <strong>the</strong> data from <strong>the</strong>se localities may'be indicative <strong>of</strong> conditions in <strong>the</strong> area in which <strong>the</strong> deep-trap-net fishery<br />

was centered.<br />

» The average positions <strong>of</strong> <strong>the</strong> tbermoelina were: last half <strong>of</strong> June, 21-33 feet; July, 67-77 feet; August, 69-80 feet; first 10 days <strong>of</strong> September,<br />

7Z-8Í feet. The <strong>the</strong>nuoolme h&d not yet formed in <strong>the</strong> first half <strong>of</strong> June; no readings were made in <strong>the</strong> area after September 10.

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