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Establecimiento de cuatro especies de Quercus en el sur de la ...

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Capítulo 4<br />

Time to emerg<strong>en</strong>ce was <strong>de</strong>termined by differ<strong>en</strong>t variables in each species<br />

(Table 2), but in most cases (3 species) it was r<strong>el</strong>ated to light avai<strong>la</strong>bility and to intrinsic<br />

traits r<strong>el</strong>ated to maternal origin (Fig. 1). Q. ilex was the species with the <strong>la</strong>rgest mean<br />

time to emerg<strong>en</strong>ce, which was only negativ<strong>el</strong>y <strong>de</strong>termined by light avai<strong>la</strong>bility. In Q.<br />

suber however, time to emerg<strong>en</strong>ce was negativ<strong>el</strong>y affected by seed mass, and also<br />

<strong>de</strong>p<strong>en</strong><strong>de</strong>d on mother tree. No differ<strong>en</strong>ces were found in mean time to emerg<strong>en</strong>ce<br />

betwe<strong>en</strong> <strong>de</strong>ciduous species (Tukey contrasts; z=0.892, P=0.807), which was on average<br />

lower than evergre<strong>en</strong>s. For Q. faginea, the species with the lowest time to emerg<strong>en</strong>ce,<br />

a complex set of parameters was involved: the interaction term betwe<strong>en</strong> light<br />

avai<strong>la</strong>bility and mother tree, the interaction betwe<strong>en</strong> soil moisture and mother tree,<br />

and a negative effect of seed mass. An interaction betwe<strong>en</strong> light avai<strong>la</strong>bility and<br />

mother tree was the only term retained in the mo<strong>de</strong>l for Q. pyr<strong>en</strong>aica, the species with<br />

the second lowest average time to emerg<strong>en</strong>ce. In the case of Q. faginea the interaction<br />

betwe<strong>en</strong> light and mother tree <strong>de</strong>termined neutral or negative effects of light<br />

avai<strong>la</strong>bility on time to emerg<strong>en</strong>ce <strong>de</strong>p<strong>en</strong>ding on the mother tree, while for Q.<br />

pyr<strong>en</strong>aica these effects were positive or negative <strong>de</strong>p<strong>en</strong>ding on the mother tree<br />

(Fig.1).<br />

Seedling emerg<strong>en</strong>ce was highest for Q. suber and Q. pyr<strong>en</strong>aica (73.5 % and<br />

71.0% respectiv<strong>el</strong>y; Table 1), and for these species it was <strong>de</strong>termined by the<br />

in<strong>de</strong>p<strong>en</strong><strong>de</strong>nt effects of seed mass and mother tree (Table 2). In both cases, seed mass<br />

had a positive influ<strong>en</strong>ce on seedling emerg<strong>en</strong>ce (Fig. 2). On the contrary, seed mass<br />

had a differ<strong>en</strong>t effect on emerg<strong>en</strong>ce of seedlings from differ<strong>en</strong>t mother trees in Q.<br />

faginea and Q. ilex, as revealed by a significant interaction betwe<strong>en</strong> mother and seed<br />

mass (Table 2). In the case of Q. faginea seed mass had a positive influ<strong>en</strong>ce on<br />

emerg<strong>en</strong>ce but with differ<strong>en</strong>t slopes for each maternal p<strong>la</strong>nt. On the contrary, for Q.<br />

ilex this r<strong>el</strong>ationship was either positive or negative <strong>de</strong>p<strong>en</strong>ding on the mother tree (Fig.<br />

2). In addition these two species showed lower rates of emerg<strong>en</strong>ce (54.2 % and 33.6 %<br />

respectiv<strong>el</strong>y; Table 1).<br />

115

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