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Establecimiento de cuatro especies de Quercus en el sur de la ...

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Reforestation with <strong>Quercus</strong> ilex L. and Q. suber L. by direct seeding and p<strong>la</strong>nting in southern Spain<br />

Seedling success (a combination of emerg<strong>en</strong>ce and <strong>sur</strong>vival) in directly see<strong>de</strong>d<br />

p<strong>la</strong>nts was about 17 % for Q. ilex and 32 % for Q. suber the first year (Table 2). Second<br />

year seedling success <strong>de</strong>creased 31.9 % in Q. ilex and 29.8 % in Q. suber (Table 2).<br />

Success was significantly affected by seed mass (F = 14.3, P < 0.001). In both species,<br />

successful seedlings had <strong>la</strong>rger acorns than the non successful, the effect being<br />

significant for Q. ilex (Fig. 3). Because seedling success was a combination of<br />

emerg<strong>en</strong>ce and <strong>sur</strong>vival, seed mass may have influ<strong>en</strong>ced either variable. Interestingly,<br />

the favourable effect of seed mass was f<strong>el</strong>t on <strong>sur</strong>vival in Q. ilex (F = 9.24, P = 0.002)<br />

but on emerg<strong>en</strong>ce in Q. suber (F = 10.66, P = 0.001)<br />

3.4<br />

Q. ilex<br />

b<br />

Q. suber<br />

3.2<br />

Seed dry mass (g)<br />

3.0<br />

2.8<br />

2.6<br />

2.4<br />

a<br />

c<br />

c<br />

2.2<br />

2.0<br />

Mean Mean±SE Mean±1.96*SE<br />

0 1<br />

Seeding success<br />

0 1<br />

Figure 3. Differ<strong>en</strong>ces in initial seed dry mass betwe<strong>en</strong> emerged and alive seedlings (1) and non‐emerged<br />

or <strong>de</strong>ad seedlings after 8 months. Differ<strong>en</strong>t letters repres<strong>en</strong>t statistically differ<strong>en</strong>t groups according to<br />

the post‐hoc Tukey test (P < 0.05).<br />

Seedling growth and morphology<br />

Morphological attributes differed betwe<strong>en</strong> species and with seedling age, both<br />

on the p<strong>la</strong>nting date (Supplem<strong>en</strong>tary information, App<strong>en</strong>dix S1) and 8 months <strong>la</strong>ter<br />

(Table 3). The change in root biomass g<strong>en</strong>erally was lower in the ol<strong>de</strong>r se<strong>el</strong>dings, and<br />

so did r<strong>el</strong>ative growth rate (RGR), specific leaf area (SLA) and leaf mass fraction (LMF)<br />

for both species (Table 3). The root/shoot ratio of the ol<strong>de</strong>r seedlings was higher,<br />

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