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Management of rice production systems to increase productivity

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analyzed by determining <strong>of</strong> genotypic variation in water uptake and how the<br />

water is used in the processes that lead <strong>to</strong> formation <strong>of</strong> grain yield (Fukai and<br />

Cooper, 1995). Once genotypic variation in resource capture or utilization<br />

efficiency (yield produced per unit nutrient uptake) is found, then<br />

physiological or morphological fac<strong>to</strong>rs that are responsible for such variation<br />

can be further examined.<br />

Of the major determining fac<strong>to</strong>rs <strong>of</strong> phenology, at least two can be<br />

ruled out in the tropics. Low temperature during the growing season <strong>of</strong><br />

upland <strong>rice</strong>, and pho<strong>to</strong>periodism. However, drought is clearly a major<br />

limiting fac<strong>to</strong>r, especially drought that develops during panicle development<br />

which delays flowering <strong>of</strong> <strong>rice</strong> in upland conditions (Inthapan and Fukai,<br />

1988; Fukai, 1999). Under upland field conditions, Lilley and Fukai (1994)<br />

showed that the magnitude <strong>of</strong> delay in flowering was associated with the<br />

severity <strong>of</strong> drought conditions. Development <strong>of</strong> the panicle is very sensitive <strong>to</strong><br />

soil water availability, and the rate <strong>of</strong> development is reduced by a small<br />

water deficit and ceases completely with the occurrence <strong>of</strong> severe drought.<br />

Drought is thus a major problem for upland <strong>rice</strong>. Yield loss due <strong>to</strong><br />

drought in upland <strong>rice</strong> was estimated at 60% in the 2002 growing season in<br />

The Gambia when rainfall declined by 40% from the 10‐year normal average<br />

<strong>of</strong> 760 mm. In fact, earlier results with upland <strong>rice</strong> <strong>production</strong> in The Gambia<br />

showed that upland <strong>rice</strong> grown under 700 mm <strong>to</strong> 1,000 mm rainfall may be<br />

subjected <strong>to</strong> moisture stress at any stage <strong>of</strong> crop growth due <strong>to</strong> periodic<br />

drought spells and uneven distribution. Drought‐related problems <strong>of</strong> crop<br />

establishment occur in many <strong>of</strong> the world’s rainfed <strong>rice</strong> growing areas<br />

(O’Toole, 1981). Drought affects plant growth primarily because <strong>of</strong> the<br />

shortage <strong>of</strong> water in the plant. Water is essential for plant growth, and<br />

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