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COST Action E 52 - vTI - Bund.de

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esearch PrIorITIes For beech IN brITaIN<br />

Over the past century a number of lines of active applied research have been pursued with regard<br />

to beech in Great Britain, both by the research branch of the Forestry Commission (now known as<br />

Forest Research) and by aca<strong>de</strong>mic researchers in university forestry <strong>de</strong>partments and government<br />

research institutes. The main areas of work have been:<br />

• Plant ecology and site affiliations – early work between the two world wars adopted the<br />

Continental European phyto-sociological and edaphic approaches, leading essentially to the<br />

three-fold floristic/edaphic classification of the British beechwoods. Leading researchers in this<br />

period were Tansley, Watt and Bourne, based at the university forestry <strong>de</strong>partments in Oxford<br />

and Cambridge (Tansley 1911, 1939, Watt 1923/25, 1931, 1934, Watt, Tansley 1930). The<br />

Chiltern and Cotswold beechwoods received particular attention. Later woodland ecologists<br />

such as Peterken and Rodwell have refined the basic classification of the British beech woodland<br />

communities, providing finer <strong>de</strong>tail (Peterken 1993, Rodwell 1991).<br />

• establishment and productivity – post Second World War research, based at the Forestry<br />

Commission’s Alice Holt Research Station, focussed on improving techniques for the establishment<br />

of new beech crops, particularly on rendzina soils/calcareous grasslands within the British<br />

natural range. There was also a programme of tree breeding, with early provenance trials, elite<br />

tree selection, progeny trials and seed orchard establishment (Savill, Fennessy, Samuel 2005).<br />

Leading Forestry Commission researchers on beech were J. M. B. Brown and D. Fourt (Brown<br />

1953). Current advice would be to establish beech crops at 2,500 stems/ha or i<strong>de</strong>ally more. Crops<br />

established at 1,100 stems/ha require intensive pruning to achieve satisfactory timber crops and<br />

this is rarely applied due to high costs for the required labour inputs.<br />

• diseases and pests – the long-standing disease of beech in Great Britain was beech bark disease,<br />

caused by an initial infestation by the felted beech coccus insect (Cryptococcus fagisuga) followed<br />

by lesion infection with the fungus Nectria coccinea. Together these cause significant canker and<br />

die-back of beech from the pole-stage onwards and were the subject of research at Alice Holt in the<br />

19 0s and 1980s (Lonsdale, Wainhouse 198 ). More recently attention has turned to the highly<br />

aggressive Phytophthora fungus-like pathogens that have become established in western parts of<br />

Great Britain (Phytophthora ramorum and Phytophthora kernoviae). Although the main hosts for<br />

these species in Great Britain are ornamental shrub species such as Rhodo<strong>de</strong>ndron, Azalea, Syringa<br />

and Pieris, these pathogens have <strong>de</strong>monstrated an ability to infect beech trees growing in close<br />

proximity, particularly in the warm-moist climates of south-western Britain. More recently Larix<br />

kaempferi has been affected in this region. A programme of research is currently being pursued,<br />

led by Forest Research at Alice Holt, to <strong>de</strong>velop avoidance and mitigation measures for these new<br />

diseases across a range of tree species. Grey squirrels remain the main mammalian pest of beech<br />

within its British native range, causing significant bark stripping damage, especially at the poletimber<br />

stage. This species was introduced to Britain in the 1800s and has since spread wi<strong>de</strong>ly,<br />

displacing the native, and less damaging, red squirrel. A variety of research and <strong>de</strong>velopment<br />

approaches has been pursued to enable effective control of grey squirrels in the forestry context<br />

by trapping and to examine the potential applicability of immuno-contraception.<br />

• drought damage and die-back – since the severe summer droughts of 19 5 – 19 6, increasing<br />

evi<strong>de</strong>nce has been found of mature beech within the British natural range suffering apparent<br />

drought damage (Peterken, Mountford 1996). This leads to crown recession and partial<br />

136

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