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Yellowstone's Northern Range - Greater Yellowstone Science ...

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Figure 7.3. Bison<br />

winter range. Since<br />

the early 1980s,<br />

bison have learned of<br />

potential winter<br />

rallges north of the<br />

park boundary,<br />

leading to a variety<br />

of controversial<br />

management actions,<br />

including public<br />

hunting ill the 1980s<br />

and capture, test,<br />

and slaughter<br />

operations more<br />

recently. Map by<br />

Yellowstolle Spatial<br />

Analysis Center alld<br />

<strong>Yellowstone</strong> Center<br />

for Resources.<br />

r c<br />

/"'\ ." ( . ..f .. '-...<br />

'. v.-J<br />

i ..<br />

" / i' .. _ ..(<br />

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i<br />

i<br />

THE NORTHERN RANGE<br />

84<br />

<strong>Northern</strong> Winter <strong>Range</strong><br />

Bison <strong>Range</strong><br />

~ 6.2 kin<br />

d f-I -(-lO-tl m<br />

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i<br />

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/<br />

J<br />

V J \<br />

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)<br />

evidence offood shortages caused by overgrazing<br />

in the park. Meagher (1989a, 1989b, 1996)<br />

evaluated these movements and concluded that<br />

much of the movements, if not all, are associated<br />

with bison learning to use hard-packed snowmobile<br />

roads for ingress and egress to the park. The<br />

grassland studies cited earlier in this document<br />

indicate that the range is not overgrazed.<br />

Meagher's observations further indicate that bison<br />

"roam" regardless of snow depth or forage availability.<br />

Once the migratory habit is learned, as it<br />

now is in the northern range bison herd and near<br />

the west boundary, the bison will quite naturally be<br />

inclined to move to lower elevations in the winter,<br />

just as the elk do. The animals shot by hunters or<br />

by management agencies after leaving the park in<br />

recent years were in good to even excellent<br />

condition as judged by body fat; they were not<br />

starving for lack of food in the park (K. Aune,<br />

Mont. Dept. Fish, Wildl. and Parks., unpubl. data;<br />

M. Meagher, U.S. Geol. Surv., pers. commun.)<br />

They were moving, as bison moved for millennia<br />

prior to the settlement of the west by Europeans.<br />

Kirkpatrick et al. (1996) compared bison<br />

pregnancy rates of the smaller northern herd with<br />

those of the more robust Mary Mountain herd.<br />

They concluded that the northern herd, which was<br />

considered to be below its ecological carrying<br />

capacity (ECC), had higher pregnancy and birth<br />

rates than bison in the Mary Mountain area, which<br />

was considered to be near or at ECC. This was<br />

broadly considered to be physiological evidence of<br />

density dependence and natural regUlation in<br />

<strong>Yellowstone</strong> bison. DelGuidice et al. (1994), while<br />

not directly addressing the natural regulation<br />

question, provided physiological evidence also<br />

suggestive of the northern bison herd being below<br />

ECC and the Mary Mountain herd being at or near<br />

ECC.<br />

Bison, unlike elk, deer, moose, bighorn<br />

sheep, and pronghorn, have few constituencies to<br />

champion their cause on public or private lands<br />

outside of <strong>Yellowstone</strong> Park. As noted above, their<br />

population has increased and they have learned

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