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Yellowstone's Northern Range - Greater Yellowstone Science ...

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THE NORTHERN RANGE<br />

Figure 7.13.<br />

NumelVus<br />

scavengers quickly<br />

consume the<br />

carcasses of elk and<br />

other ungulates on<br />

the northern range.<br />

NPSphoto.<br />

Figure 7.14.<br />

<strong>Yellowstone</strong> grizzly<br />

bears are adaptable<br />

omnivores, adjusting<br />

to the varying<br />

supplies of preferred<br />

foods. Scat analysis<br />

has shown that in<br />

years of poor<br />

whitehark pine lIut<br />

crops, hears<br />

compensated by<br />

increasing their<br />

consumption of<br />

ungulates. The<br />

northern <strong>Yellowstone</strong><br />

elk herd is an<br />

important component<br />

of the diet of the<br />

threatened grizzly<br />

bem:<br />

Interagency Grizzly<br />

Bear Study Team.<br />

100<br />

98<br />

ecological and ungulate<br />

management issue has entered<br />

an imp0l1ant new phase<br />

(Phillips and Smith 1996,<br />

Schullery 1996). This addition<br />

to the suite of predator/<br />

scavengers (grizzly and bl,ack<br />

bears, mountain lions, wolverines,<br />

coyotes, faxes, plus<br />

numerous bird species) is<br />

significant (Figure 7.15).<br />

Several writers have pointed<br />

out the incompleteness of the<br />

natural regulation experiment<br />

as long as the full complement<br />

of native mammalian predators<br />

UNG =0.518 +0.065 YR- 0.0165 PIAL<br />

r' was not in place (Peek 1980,<br />

=0.99<br />

R<br />

Houston 1982, Mech 1991).<br />

The arrival of the wolf is<br />

Ungulates<br />

perhaps the single most<br />

important event in the evolution<br />

of northern range manage­<br />

, ,<br />

, " ,, ,<br />

ment since the 1967 Senate<br />

,<br />

,,<br />

, hearings that led to the<br />

,,<br />

,<br />

cessation of traditional elk,<br />

Q"""""<br />

, Whitcbark Pine Nuts<br />

bison, and pronghorn control.<br />

,,<br />

, "'~--------Q.. ...---........."9<br />

Prior to wolf reintroduction,<br />

North American wolf<br />

YEAR<br />

experts, while expressing a<br />

majority opinion about the<br />

probable effects of <strong>Yellowstone</strong><br />

°7~7-------7LI8-------7~~-------8~~~----~8~1------~8L2------~83<br />

Recent calculations by Singer et al. (1997) suggest<br />

each grizzly bear within the calving range of the<br />

northern elk herd kills and consumes, on the<br />

average, about 15 elk calves per year. Of course,<br />

this rate would vary from bear to bear since some<br />

animals specialize more in the hunting of elk<br />

calves than others, and the rate would also vary<br />

from year to year (Singer et al. 1997). Based on<br />

close observations of a small number of bears for<br />

short time periods, French and French (1990)<br />

calculated that some bears killed at least one calf<br />

per day, on average, during the four to six weeks<br />

each year that calves are vulnerable to bear<br />

predation.<br />

With the reintroduction of wolves to the<br />

northern range in 1995, the entire northern range<br />

wolves, also had minority views that expressed a<br />

wide range of opinion in the probable effects (Vales<br />

and Peek 1990, Boyce and Gaillard 1992, Singer<br />

1991, Lyme et aI. 1993). All predicted some<br />

decline in elk numbers and some predicted declines<br />

or increases in other ungulates. Singer (U.S. Geo!.<br />

Surv., unpub!' data) predicted a decline in coyotes<br />

and an increase in faxes, and no effects on grizzly<br />

and black bears, mountain lions, and wolverines.<br />

They predicted an increase in pronghorns, moose,<br />

bighorns, and beaver, and noted that mule deer<br />

should be monitored closely. They also concluded<br />

that the moderate wolf densities they expect in<br />

future, and their effects on the large herbivore<br />

populations would not likely result in any significant<br />

effects on the northern range's vegetation. In

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