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Yellowstone's Northern Range - Greater Yellowstone Science ...

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Figure 4.2. Willow<br />

distribution in<br />

<strong>Yellowstone</strong> National<br />

Park. The largest<br />

concentrations of<br />

willow are south of<br />

<strong>Yellowstone</strong> Lake<br />

along the upper<br />

<strong>Yellowstone</strong> Rivel; in<br />

the southwest comer<br />

of the park, and in<br />

tributaries of the<br />

Madison River near<br />

the west boundmy.<br />

Map by the<br />

<strong>Yellowstone</strong> Spatial<br />

Analysis Center and<br />

the <strong>Yellowstone</strong><br />

Center for<br />

Resources.<br />

has led to predictions of decreases in coyote<br />

numbers (R. Crabtree, <strong>Yellowstone</strong> Ecosystem<br />

Stud., pers. commun.), which may permit an<br />

increase in pronghorn (Berger 1991), the ungulate<br />

species most dependent upon sagebrush, and<br />

currently regarded as a species of special concern<br />

on the northern range (see pronghorn research<br />

recommendations, Chapter Seven). Further<br />

research into the ecology of the various sagebrush<br />

species across the northern winter range is needed,<br />

especially into the factors contributing to the<br />

decline of the Wyoming SUbspecies, and the<br />

apparent thriving of the basin subspecies. The role<br />

of pronghorn in the vitality of sagebrush and<br />

climate change appears crucial. Continued<br />

monitoring of all of these interactive forces in the<br />

BLA seems warranted.<br />

WILLOWS<br />

About one-half of one percent (0.54 percent)<br />

of the <strong>Yellowstone</strong> Park's willow<br />

communities are on the northern<br />

range (Figure 4.2). The greatest<br />

concentrations of willows in<br />

<strong>Yellowstone</strong> National Park are<br />

south of <strong>Yellowstone</strong> Lake, the<br />

southwest corner of the park in<br />

the Bechler River area, and near<br />

the west boundary north of the<br />

Madison River. Willow abundance<br />

in the park is almost<br />

entirely defined by elevation and<br />

precipitation; areas that are<br />

above 7,000 feet and have more<br />

than 20 inches of annual precipitation<br />

comprise 74 percent of the<br />

park's willow communities, and<br />

areas that are above 7,000 feet<br />

andlor have more than 20 inches<br />

of annual precipitation comprise<br />

more than 99 percent of the<br />

park's willow communities.<br />

These are high-elevation sites,<br />

not on the winter ranges of<br />

ungulates, except possibly that of<br />

moose.<br />

THE NORTHERN RANGE<br />

~<br />

1<br />

1<br />

46<br />

Willows have been described as an "insignificant<br />

component" of the <strong>Yellowstone</strong> area for at<br />

least the past 20,000 years (Mullenders and<br />

Coremans 1996, Mullenders et aJ. 1996). That<br />

should be taken to mean that they occupy comparatively<br />

little area. It should not be taken to mean<br />

that they do not have significant ecological effects<br />

on northern range plant and animal communities, a<br />

subject dealt with later in this section.<br />

Declines in willows were noted early in the<br />

northern range's written history. Houston, based<br />

on comparison of historic photographs with<br />

modern retakes, estimated that during the past<br />

century, willows may have declined in area from<br />

about 0.8 percent of the northern range (that is,<br />

slightly less than 1 percent) to about 0.4 percent<br />

(that is, less than half of I percent). Houston<br />

(1982) and Chadde and Kay (1991) published<br />

photographic evidence of heavier willow growth<br />

along some park streams prior to 1900.<br />

There are approximately 24 species of<br />

c=J <strong>Northern</strong> Winter <strong>Range</strong><br />

_<br />

Willow Habitat<br />

I 6.2km I<br />

(10mi)<br />

!<br />

.. _.,_ .. _ .. __ .. r.<br />

N<br />

'1<br />

)<br />

\..<br />

"';.<br />

./ ..<br />

r-"';---<br />

,,..r"<br />

'l..f:'<br />

1<br />

(<br />

1 ( l.<br />

: ~ "1<br />

1 , ~ :<br />

i \ . 1<br />

~ .. -->.C._ .. _.~_ .. _ .. _ .. _ -.1.-1

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