Yellowstone's Northern Range - Greater Yellowstone Science ...
Yellowstone's Northern Range - Greater Yellowstone Science ...
Yellowstone's Northern Range - Greater Yellowstone Science ...
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Figure 4.2. Willow<br />
distribution in<br />
<strong>Yellowstone</strong> National<br />
Park. The largest<br />
concentrations of<br />
willow are south of<br />
<strong>Yellowstone</strong> Lake<br />
along the upper<br />
<strong>Yellowstone</strong> Rivel; in<br />
the southwest comer<br />
of the park, and in<br />
tributaries of the<br />
Madison River near<br />
the west boundmy.<br />
Map by the<br />
<strong>Yellowstone</strong> Spatial<br />
Analysis Center and<br />
the <strong>Yellowstone</strong><br />
Center for<br />
Resources.<br />
has led to predictions of decreases in coyote<br />
numbers (R. Crabtree, <strong>Yellowstone</strong> Ecosystem<br />
Stud., pers. commun.), which may permit an<br />
increase in pronghorn (Berger 1991), the ungulate<br />
species most dependent upon sagebrush, and<br />
currently regarded as a species of special concern<br />
on the northern range (see pronghorn research<br />
recommendations, Chapter Seven). Further<br />
research into the ecology of the various sagebrush<br />
species across the northern winter range is needed,<br />
especially into the factors contributing to the<br />
decline of the Wyoming SUbspecies, and the<br />
apparent thriving of the basin subspecies. The role<br />
of pronghorn in the vitality of sagebrush and<br />
climate change appears crucial. Continued<br />
monitoring of all of these interactive forces in the<br />
BLA seems warranted.<br />
WILLOWS<br />
About one-half of one percent (0.54 percent)<br />
of the <strong>Yellowstone</strong> Park's willow<br />
communities are on the northern<br />
range (Figure 4.2). The greatest<br />
concentrations of willows in<br />
<strong>Yellowstone</strong> National Park are<br />
south of <strong>Yellowstone</strong> Lake, the<br />
southwest corner of the park in<br />
the Bechler River area, and near<br />
the west boundary north of the<br />
Madison River. Willow abundance<br />
in the park is almost<br />
entirely defined by elevation and<br />
precipitation; areas that are<br />
above 7,000 feet and have more<br />
than 20 inches of annual precipitation<br />
comprise 74 percent of the<br />
park's willow communities, and<br />
areas that are above 7,000 feet<br />
andlor have more than 20 inches<br />
of annual precipitation comprise<br />
more than 99 percent of the<br />
park's willow communities.<br />
These are high-elevation sites,<br />
not on the winter ranges of<br />
ungulates, except possibly that of<br />
moose.<br />
THE NORTHERN RANGE<br />
~<br />
1<br />
1<br />
46<br />
Willows have been described as an "insignificant<br />
component" of the <strong>Yellowstone</strong> area for at<br />
least the past 20,000 years (Mullenders and<br />
Coremans 1996, Mullenders et aJ. 1996). That<br />
should be taken to mean that they occupy comparatively<br />
little area. It should not be taken to mean<br />
that they do not have significant ecological effects<br />
on northern range plant and animal communities, a<br />
subject dealt with later in this section.<br />
Declines in willows were noted early in the<br />
northern range's written history. Houston, based<br />
on comparison of historic photographs with<br />
modern retakes, estimated that during the past<br />
century, willows may have declined in area from<br />
about 0.8 percent of the northern range (that is,<br />
slightly less than 1 percent) to about 0.4 percent<br />
(that is, less than half of I percent). Houston<br />
(1982) and Chadde and Kay (1991) published<br />
photographic evidence of heavier willow growth<br />
along some park streams prior to 1900.<br />
There are approximately 24 species of<br />
c=J <strong>Northern</strong> Winter <strong>Range</strong><br />
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