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moisture affects wheat development is believed<br />
to depend upon the stage of growth at the<br />
time it occurs. Reported effects include a reduction<br />
in the number of tillers, the fertility of<br />
the spikes, and the size of the grain.<br />
The effects of flooding on wheat development<br />
have been studied at the Northwest<br />
Agricultural Research Center (CIANO) Mexico,<br />
during the past two seasons. The soils in this<br />
region are heavy clays and farmers have an<br />
ample supply of low cost irrigation water;<br />
these conditions suggested that an excess of<br />
soil moisture after irrigating might be a yieldlimiting<br />
factor.<br />
In 1969-70, wheat production in normally<br />
irrigated plots was compared with that in plots<br />
flooded for 4 or 8 days at different times during<br />
the growing season. The soil was a heavy<br />
clay with a bulk density that varied with depth<br />
from 1.1 in the 0-30 cm horizon, to 1.35 in the<br />
60-90 cm horizon, to 1.20 at the 120 cm depth.<br />
Apparently the more dense horizon below 60 cm<br />
was almost impermeable to water, as little<br />
penetration occurred below that depth, even<br />
in plots flooded for as long as 4 days. Oxygen<br />
diffusion rates were measured as soon as possible<br />
after flooding and the measurements<br />
were continued until the rates were adequate<br />
for normal growth. After a normal irrigation,<br />
the oxygen diffusion rate remained below 0.2<br />
micrograms/cm 2 /min at a soil depth of 15<br />
cm for a period of 9 to 12 days. Following<br />
4 days of flooding, the oxygen diffusion rate<br />
was less than 0.2 micrograms/cm 2 /min at a<br />
15 cm depth for 12 to 15 days.<br />
Flooding was found to affect plant development<br />
in several ways:<br />
Flooding for 4 days at planting time reduced<br />
plant emergence by 17 percent and plant<br />
height at 26 days by 11 percent. The number<br />
of heads .and grain yield decreased very<br />
slightly.<br />
Flooding for 4 days, at 15 and 30 days<br />
after planting, increased plant height and the<br />
number of tillers per plant at 53 days. Straw<br />
yields were increased by about 16 percent.<br />
At 45 days after planting, flooding reduced<br />
plant height, tiller density at 53 days, grain<br />
size, grain yield and straw yield; it did not<br />
change the number of grains per head.<br />
Flooding for 4 days at 60 and 75 days after<br />
planting had no significant effect on wheat<br />
development.<br />
Flooding for 8 days at 65 days after planting<br />
slightly increased both grain size and<br />
yield.<br />
The INIA variety used in this study flowered<br />
at 61 days after planting.<br />
Evidently, as far as grain yield was concerned,<br />
the INIA wheat variety was most sensitive<br />
to a low oxygen diffusion rate at planting<br />
time and about 45 days later. The effect<br />
of flooding at 45 days was similar to that<br />
observed in 1968-69, when flooding at 38 days<br />
reduced both plant height and straw yields.<br />
Surprisingly, flooding for four days at 15, 30,<br />
60 and 75 days after planting, did not reduce<br />
grain yields; while flooding for 8 days, at 65<br />
days after planting, slightly increased yields.<br />
Similar results were obtained in a growth<br />
chamber study at the University of California<br />
at Riverside, which found that maximum production<br />
of wheat grain occurred at atmospheric<br />
oxygen levels about one-fourth normal<br />
level.<br />
Results obtained in studies conducted during<br />
the past two years clearly indicate that<br />
present wheat varieties can tolerate several<br />
days of flooding without important reductions<br />
in yield. Further information is needed on the<br />
quantitative effects of low oxygen diffusion<br />
rates occurring about 45 days after planting.<br />
Timing of the Second Irrigation for Wheat<br />
Field studies conducted in India over the<br />
past 5 years have shown that maximum wheat<br />
yields are obtained when the first irrigation<br />
after planting is made 20 to 25 days following<br />
planting, when crown root development<br />
begins. CIANO experiments in 1969-1970 have<br />
obtained more information on this point. Second<br />
irrigations in both years were applied 15,<br />
22, 29, 36 and 43 days after planting. The varieties<br />
used in the 1969 experiments were Sonora<br />
64 and Tobari, while Sonora 64 and INIA<br />
were used in 1970.<br />
Grain yields of Sonora 64 and Tobari were<br />
5.00 and 4.78 ton/ha, respectively, in 1969.<br />
Timing of the second irrigation did not affect<br />
yields significantly.<br />
The INIA variety produced equal yields of<br />
3.67 ton/ha in 1970 when the second irrigation<br />
was applied 15, 22, 29, and 36 days after<br />
planting. Delaying the second irrigation until<br />
43 days after planting reduced the grain yield<br />
to 3.46 ton/ha.<br />
The Sonora 64 variety produced the highest<br />
yields, 4.30 ton/ha, when the second irrigation<br />
was applied at 22 or 29 days after planting.<br />
Yields were slightly less (4.00 ton/ha) when<br />
the second irrigation was applied at 15 or 36<br />
days after planting.<br />
These results are not conclusive, but, in<br />
the case of Sonora 64, do provide some evidence<br />
that application of the second irrigation<br />
at about 25 days after planting is essential<br />
for maximum grain production.<br />
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