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moisture affects wheat development is believed<br />

to depend upon the stage of growth at the<br />

time it occurs. Reported effects include a reduction<br />

in the number of tillers, the fertility of<br />

the spikes, and the size of the grain.<br />

The effects of flooding on wheat development<br />

have been studied at the Northwest<br />

Agricultural Research Center (CIANO) Mexico,<br />

during the past two seasons. The soils in this<br />

region are heavy clays and farmers have an<br />

ample supply of low cost irrigation water;<br />

these conditions suggested that an excess of<br />

soil moisture after irrigating might be a yieldlimiting<br />

factor.<br />

In 1969-70, wheat production in normally<br />

irrigated plots was compared with that in plots<br />

flooded for 4 or 8 days at different times during<br />

the growing season. The soil was a heavy<br />

clay with a bulk density that varied with depth<br />

from 1.1 in the 0-30 cm horizon, to 1.35 in the<br />

60-90 cm horizon, to 1.20 at the 120 cm depth.<br />

Apparently the more dense horizon below 60 cm<br />

was almost impermeable to water, as little<br />

penetration occurred below that depth, even<br />

in plots flooded for as long as 4 days. Oxygen<br />

diffusion rates were measured as soon as possible<br />

after flooding and the measurements<br />

were continued until the rates were adequate<br />

for normal growth. After a normal irrigation,<br />

the oxygen diffusion rate remained below 0.2<br />

micrograms/cm 2 /min at a soil depth of 15<br />

cm for a period of 9 to 12 days. Following<br />

4 days of flooding, the oxygen diffusion rate<br />

was less than 0.2 micrograms/cm 2 /min at a<br />

15 cm depth for 12 to 15 days.<br />

Flooding was found to affect plant development<br />

in several ways:<br />

Flooding for 4 days at planting time reduced<br />

plant emergence by 17 percent and plant<br />

height at 26 days by 11 percent. The number<br />

of heads .and grain yield decreased very<br />

slightly.<br />

Flooding for 4 days, at 15 and 30 days<br />

after planting, increased plant height and the<br />

number of tillers per plant at 53 days. Straw<br />

yields were increased by about 16 percent.<br />

At 45 days after planting, flooding reduced<br />

plant height, tiller density at 53 days, grain<br />

size, grain yield and straw yield; it did not<br />

change the number of grains per head.<br />

Flooding for 4 days at 60 and 75 days after<br />

planting had no significant effect on wheat<br />

development.<br />

Flooding for 8 days at 65 days after planting<br />

slightly increased both grain size and<br />

yield.<br />

The INIA variety used in this study flowered<br />

at 61 days after planting.<br />

Evidently, as far as grain yield was concerned,<br />

the INIA wheat variety was most sensitive<br />

to a low oxygen diffusion rate at planting<br />

time and about 45 days later. The effect<br />

of flooding at 45 days was similar to that<br />

observed in 1968-69, when flooding at 38 days<br />

reduced both plant height and straw yields.<br />

Surprisingly, flooding for four days at 15, 30,<br />

60 and 75 days after planting, did not reduce<br />

grain yields; while flooding for 8 days, at 65<br />

days after planting, slightly increased yields.<br />

Similar results were obtained in a growth<br />

chamber study at the University of California<br />

at Riverside, which found that maximum production<br />

of wheat grain occurred at atmospheric<br />

oxygen levels about one-fourth normal<br />

level.<br />

Results obtained in studies conducted during<br />

the past two years clearly indicate that<br />

present wheat varieties can tolerate several<br />

days of flooding without important reductions<br />

in yield. Further information is needed on the<br />

quantitative effects of low oxygen diffusion<br />

rates occurring about 45 days after planting.<br />

Timing of the Second Irrigation for Wheat<br />

Field studies conducted in India over the<br />

past 5 years have shown that maximum wheat<br />

yields are obtained when the first irrigation<br />

after planting is made 20 to 25 days following<br />

planting, when crown root development<br />

begins. CIANO experiments in 1969-1970 have<br />

obtained more information on this point. Second<br />

irrigations in both years were applied 15,<br />

22, 29, 36 and 43 days after planting. The varieties<br />

used in the 1969 experiments were Sonora<br />

64 and Tobari, while Sonora 64 and INIA<br />

were used in 1970.<br />

Grain yields of Sonora 64 and Tobari were<br />

5.00 and 4.78 ton/ha, respectively, in 1969.<br />

Timing of the second irrigation did not affect<br />

yields significantly.<br />

The INIA variety produced equal yields of<br />

3.67 ton/ha in 1970 when the second irrigation<br />

was applied 15, 22, 29, and 36 days after<br />

planting. Delaying the second irrigation until<br />

43 days after planting reduced the grain yield<br />

to 3.46 ton/ha.<br />

The Sonora 64 variety produced the highest<br />

yields, 4.30 ton/ha, when the second irrigation<br />

was applied at 22 or 29 days after planting.<br />

Yields were slightly less (4.00 ton/ha) when<br />

the second irrigation was applied at 15 or 36<br />

days after planting.<br />

These results are not conclusive, but, in<br />

the case of Sonora 64, do provide some evidence<br />

that application of the second irrigation<br />

at about 25 days after planting is essential<br />

for maximum grain production.<br />

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