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During the past year, grain-to-stover ratios<br />

were used in: (1) a mass selection scheme,<br />

and (2) selection among and within 51 lines.<br />

The germplasm used in the mass selection<br />

study was a composite derived from a cross<br />

between a short early corn, from Antigua in<br />

the Caribbean and a tall Tuxpeno from Mexico.<br />

A program of mass selection in this composite<br />

was started in Poza Rica, Veracruz in<br />

1969, using a selecton intensity of five percent<br />

for plants with the highest grain-to-stover ratio.<br />

The population has undergone three cycles of<br />

mass selection. During the second cycle, some<br />

selection pressure was directed toward lowering<br />

ear height, selecting only those plants with<br />

the uppermost ear below 161 em. for the<br />

grain-to-stover determinations. Badly diseased<br />

or otherwise undesirable plants were de-tasseled<br />

during the third cycle of mass selection.<br />

Progress from the first cycle of selection<br />

was evaluated in the summer of 1969 in a<br />

performance trial with ten replicates. This trial<br />

included: the original population Co; the first<br />

cycle population, C I (Me) from the grain stover<br />

program; and a first cycle population derived<br />

from selection for grain weight per plant C l<br />

(M).<br />

Three plant densities (21,739; 43,478; and<br />

65,217 plants/ha) were used in a split-plot<br />

design. Data were recorded on days-to-silking,<br />

plant and ear height, day-to-pollen shedding,<br />

number of ears-per-100 plants, stover weight,<br />

grain weight, shelling percentage, and percent<br />

of lodged plants.<br />

Mean values and the realized gain for<br />

grain-to-stover ratio and grain yield of the<br />

populations are presented in Table 1. Relative<br />

to the original population, one cycle of<br />

selection resulted in a gain of 2.9 percent in<br />

grain-to-stover ratio at the rate of planting<br />

similar to the one used during selection. Negative<br />

values were obtained when the population<br />

was evaluated at higher plant densities.<br />

The population resulting from selection for<br />

grain yield per plant showed no gain in grainto-stover<br />

ratio at all three plant densities.<br />

It is worth noting that the gains in yield<br />

after one cycle of selection for grain-to-stover<br />

ratio lC I (Me) vs. Co] were 2.2 percent, 2.4<br />

percent, and 6.3 percent at 21,739; 43,478; and<br />

62,517 plants per hectare, respectively. This<br />

seems to indicate that selection for grain-tostover<br />

ratio resulted in considerable improvement<br />

in yield and tolerance to high plant densities.<br />

One cycle of mass selection based on<br />

yield-per-plant lC I (M) vs. Co] produced a<br />

gain of 3.2 percent in grain yield at 21,739<br />

plants per hectare, but there was no gain at<br />

43,478 plants/ha, and a reduction of 4.9 percent<br />

at 65,217 plants/ha.<br />

There were changes associated with the<br />

selection for grain-to-stover ratio in other<br />

agronomic characteristics, also. In general,<br />

the population C I (Me) was earlier, with shorter<br />

plants and lower ears, and better standability<br />

than the original population.<br />

The outstanding feature of the selection for<br />

grain-to-stover ratio seems to be the correlated<br />

responses which enable the resulting<br />

population to be: (1) more productive, (2) more<br />

tolerant to high plant densities, (3) earlier, and<br />

(4) shorter.<br />

Estimation of Heritability and Correlations<br />

with Grain·to-Stover Ratio<br />

Using fifty families showing the highest<br />

grain-to-stover ratio from a mass selection<br />

program for this character, an experiment was<br />

planted in Poza Rica, Veracruz, Mexico, to<br />

estimate the heritability of grain-to-stover ratio<br />

by parent-offspring regression and to compare<br />

simple correlations between grain-to-stover<br />

ratio and eight other characters. These characters<br />

were: plant height, ear height, day-topollen<br />

shedding, days-to-silking, lodged plants,<br />

number of ears, stover weight, and grain<br />

weight.<br />

TABLE 13. Mean values and % realized gain for<br />

grain/stover ratio and grain yield of three Compo<br />

C populations at three planting densities. Poza<br />

Rica, Veracruz, Mexico. 1969.<br />

Grain/stover<br />

Grain yield (kg/ha)<br />

Mean % Gain Mean % Gain<br />

21,739 pl/ha<br />

Co 0.68 0.0 3391 0.0<br />

Cl (Me) 0.70 + 2.9 3467 + 2.2<br />

Cl (M) 0.67 - 1.5 3500 + 3.2<br />

43,478 pl/ha<br />

Co 0.68 0.0 5022 0.0<br />

Cl (Me) 0.67 - 1.5 5141 + 2.4<br />

Cl (M) 0.67 - 1.5 5022 0.0<br />

65,217 pi/he<br />

Co 0.64 0.0 5369 0.0<br />

Cl (Me) 0.60 - 6.2 5706 + 6.3<br />

Cl (M) 0.60 - 6.2 5109 - 4.9<br />

The estimated heritability for grain-to-stover<br />

ratio was 0.16 -+- 0.27, which falls within the<br />

same range as grain yield. This value indicates<br />

that selection for this charac'ter must necessary<br />

show relatively small gains per cycle.<br />

The correlation coefficients are presented<br />

in Table 2A. Highly significant negative', correlations<br />

were found between grain-to-stoVer<br />

26

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