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sound scientific approach, incorporate and<br />

utilize it in the breeding program.<br />

Each season several hundred crosses are<br />

made between superior <strong>CIMMYT</strong> dwarf phenotypes<br />

and materials from the International<br />

Spring Wheat Rust Nurseries (ISWRN) which<br />

have been found to be not only resistant in<br />

Mexico but also in several other parts of the<br />

world. Mexico, and especially the summer<br />

nursery site in Toluca, is ideal for screening<br />

for stem rust and stripe rust and to a lesser<br />

degree for leaf rust resistance. Although leaf<br />

and glume blotch Septoria tritiei and Septoria<br />

nodorum occur in Mexico, especially in the<br />

summer Toluca nursery, it seldom reaches<br />

epiphitotic proportions. Consequently, it is impossible<br />

to select effectively for res'istance<br />

under Mexican conditions. Other diseases,<br />

such as scab(Fusarium or Giberella head-blight)<br />

and powdery mildew (Erisyphe graminis var.<br />

tritiei) are rare and never epidemic. In these<br />

cases too it is necessary to bring heavy selection<br />

pressure to bear on populations segregating<br />

for resistance if resistant varieties are to<br />

be ~eveloped. However, <strong>CIMMYT</strong> being an<br />

international research institute, and directly<br />

or indirectly cooperating with many countries,<br />

is in a strong position to assist in the development<br />

of lines or varieties combining resistance<br />

to all of the aforementioned diseases.<br />

Materials reported as being resistant to Septoria,<br />

scab and mildew in national programs in<br />

other parts of the world are crossed widely<br />

with <strong>CIMMYT</strong> dwarfs in Mexico. The F 2 bulk<br />

materials derived from these crosses are sent<br />

for screening to cooperating countries in other<br />

parts of the world. Seed of any outstanding<br />

lines identified through these procedures again<br />

are sent back to <strong>CIMMYT</strong> for further crossing<br />

and up-grading of the gene pool.<br />

Within the past year a <strong>CIMMYT</strong> Rust Research<br />

Laboratory, under the direction of Dr.<br />

S. Rajaram, has been established at Chapingo.<br />

Research at this laboratory is done to identify<br />

the genes for resistance in basic and advanced<br />

generation lines· of bread and durum wheats<br />

and in triticales. The data supplement those<br />

from the field. The summarized data from the<br />

past year appear below.<br />

Identification of Races: Three hundred (300)<br />

collections of stem rust were made from<br />

Sonora, Toluca, Chapingo, Roque and Huamantia<br />

areas during 1969 and these were identified<br />

on the set of International Differentials.<br />

The most prevalent major races found were 11,<br />

12, 15, 113 and 151. These races are further<br />

defined in terms of their pathogenecity genes<br />

on genetic stocks.<br />

In 1969, 112 genetic wheat stocks were received<br />

from Dr. I. A. Watson (Australia) for use<br />

in analyzing the Mexican stem rust flora as<br />

a part of the International Virulence Gene<br />

Survey. The results have given a good idea<br />

about the effectiveness of the various stem<br />

rust genes. On the basis of these tests 7 lines<br />

with different genes are now used as supplemental<br />

rust differentials. These genes are internationally<br />

used in various breeding programs,<br />

and listed as follows:<br />

1) Sr 6 (Me Marachy); 2) Sr 11 (Yalta); 3) Sr 9b<br />

(Gamenya); 4) Sr n. (Mengavi); 5) Sr 8 (Mentana);<br />

6) Sr 17 (Selkirk), and 7) Yuma.<br />

The definition of the pathogenecity of the<br />

standard races, in terms of precise genes has<br />

permitted a more conclusive and realistic analysis<br />

of the <strong>CIMMYT</strong> breeding materials. It<br />

has also made possible the detection of new<br />

genes in the program.<br />

Based on the analysis of stem rust races<br />

from last summer's collection in Toluca, Chapingo,<br />

Roque and Huamantla it is clear that<br />

no single gene would give very effective protection.<br />

The summarized results are given in<br />

Table W6.<br />

Six strains of stem rust having different<br />

combinations of genes for pathogenecity were<br />

TABLE W6. Analysis of the total stem rust collection<br />

made from Toluca and Chapingo, Mexico,<br />

during the summer of 1969, showing virulence in<br />

relation to different tester genes, Mexican cultivars<br />

and other resistant genotypes.<br />

Me Murechy<br />

V.It.<br />

G.meny.<br />

Mengevi<br />

Ment.n.<br />

Siete Cerros<br />

Pitlc 62<br />

Sonor.64<br />

Tobarl66<br />

Nor_Ie 66<br />

Nortefio 67<br />

Selkirk<br />

Vuma<br />

W.ldron<br />

Chris<br />

Crim<br />

Genotype<br />

Sr6<br />

Sr11<br />

Sr9b<br />

SrTt<br />

SrI<br />

(Sr6 + Sr11)<br />

(Sr11 +)<br />

(Sr11 +)<br />

(SrI + Sr9b +)<br />

(Sr6 + Sr17)<br />

Percent.ges of<br />

Isol.tes Sh_ing<br />

Virulence<br />

(on seedlings)<br />

57<br />

51<br />

66<br />

33<br />

11<br />

32<br />

71<br />

22<br />

1<br />

o<br />

1<br />

o<br />

44<br />

22<br />

65

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