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To generate a maize population of maximum<br />

variability, <strong>CIMMYT</strong> has begun to put<br />

together, into a single composite, seed from<br />

5,000 of the more than 10,000 collections<br />

available in the germplasm bank in Mexico.<br />

Six composites were made by taking 15 seeds<br />

of each collection. These composites were<br />

planted at ten-day intervals in a neutral environment<br />

at Tlaltizapan in the State of Morelos<br />

Plants of the third and fourth sowing date~<br />

were detasseled. Natural selection was important<br />

in eliminating some of the collections<br />

insofar as seed production was concerned but<br />

some plants, which did not produce seed,<br />

probably produced pollen which was represented<br />

in the seed harvested. The extent of<br />

differential loss is not subject to determination.<br />

Reaction to soil temperature and climate<br />

probably resulted in variable elimination of<br />

several of the collections, also.<br />

Approximately 20,300 ears of the de-tasseled<br />

rows from this first cycle of synthesis<br />

were used to prepare two balance composites.<br />

One composite was formed by taking ten<br />

seeds from each ear harvested. Twenty seeds<br />

were taken for the second composite.<br />

The first composite was subdivided by taking<br />

87,000 seeds to plant the male rows at<br />

three different sowing dates for the second<br />

cycle of intercrossing during the winter of<br />

1969-1970. The remaining 116,000 seeds were<br />

LJse to plant the female rows, which were<br />

de-tasseled. This arrangement permitted maximum<br />

recombination among early, intermediate,<br />

and 'late genotypes.<br />

The second composite will be developed<br />

wHh the remaining collections. Then, the two<br />

resulting composites will be planted together<br />

to form the complete "world composite". Following<br />

t~is compositing, the material will be<br />

divided, and recombination and selection will<br />

be continued at low, intermediate and high<br />

elevations.<br />

All potentially useful genes and/or gene<br />

combinations are not likely to be present in<br />

any single gene pool. Hence it is unlikely that<br />

such a utopian population will be created<br />

through the formation of a "world composite".<br />

It is likely, however, that a far broader mixture<br />

of genetic materials is being made than has<br />

existed before, and this may be of considerable<br />

value to maize improvement efforts everywhere.<br />

Plant Height Studies<br />

Most tropical corn plants grow too tall and<br />

are vulnerable to lodging. Even moderate<br />

winds exert tremendous force on very tall<br />

plants. In order to maintain plant standability,<br />

and to consider higher fertilizer levels aimed<br />

at higher production, it appears that varieties<br />

with shorter plants must be developed. Two<br />

separate avenues of approach are being followed<br />

to achieve shorter plants. Genetic<br />

dwarfs are being studied as one possibility,<br />

while another attempt is through recurrent selection<br />

of ear-to-row plantings and hand pollination<br />

among selected rows.<br />

The genetic dwarfs being used are brachytic<br />

1, brachytic 2, brachytic 3, anther ear,<br />

d] short, d 1 tall, and pygmy. Others, as yet<br />

not identified, are being studied. The brachytic<br />

2 has been used to convert several diverse<br />

germplasm pools to dwarf plants. Modified<br />

ear-to-row mass selection for yield is being<br />

carried out in Tuxpeno Crema I-brachytic 2.<br />

Problems of irregularity of plant height, husk<br />

cover, ear position, and other difficulties remain,<br />

but the materials appear promising. The<br />

other dwarf populations have not progressed<br />

as far, but some show real promise.<br />

<strong>CIMMYT</strong> has begun to put together, into a single<br />

composite, seed from 5,000 of the more than<br />

10,000 collections available in the germ plasm<br />

bank in Mexico. The aim is to generate a maize<br />

population of maximum variability... a "World<br />

Composite".<br />

24

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