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A Comparisonof of Two Methods<br />

of of Stratified Mass Selection<br />

Stratified Mass Selection as as usually practiced<br />

by by maize breeders involves the time<br />

consuming operation of of physically weighing<br />

the dry grain producedon on eachof of the selected<br />

plants.<br />

Selection for prolificacy has been suggested<br />

as as an an effective alternative.<br />

The study<br />

described below attempts to to compare the effectiveness<br />

of of these two systems.<br />

Two populations adapted to to the inter­<br />

inter~<br />

mediate altitudes of of Mexico (Composite II II<br />

Celaya and Puebla GroupI) I) were subjectedtoto<br />

three cycles of of Stratified Mass Selection by by<br />

grain weight and by prolificacy.<br />

The resulting<br />

populations were studied in in yield trial for a<br />

period of of three year and the data are summarized<br />

in in Table 17.<br />

It It is is apparent that the two methods were<br />

substantially equal in in their effectivenessinin increasing<br />

yield and that the yields after two<br />

and three cycles in in Composite Celaya II II<br />

and<br />

Puebla Group I, I, respectively, were equal to to<br />

or or better than the best adapted hybrid for the<br />

region. It It should be be noted, however, that the<br />

Puebla GroupI, I, Cycle22 population, could not<br />

be be recommended to to farmers because of of its its<br />

lodging susceptibility even thoughitit ly ly outyields the recommended hybrids.<br />

Another fact is is worth noting. These data<br />

do not show the clearcut tendency for the accompanying<br />

undesirable side effects -- in­<br />

increaseinin<br />

daystoto flower norinin percent lodging<br />

apparent­<br />

as as was reportedinin the studyof of Stratified Mass<br />

Selection versus Ear-to-row selection in in the<br />

two Chalquerio populations.<br />

GENETICS AND CYTOLOGY<br />

Studieson on the Chromosome<br />

Morphologyof of Maize<br />

Extensive data on on chromosome knobs of of<br />

maize have been accumulated during the last<br />

decade, beginning with the workof of Dr. Barbara<br />

McClintockinin 1958.<br />

These studies have been<br />

extremely useful in in determining the origin,<br />

relationships and migration of of the different<br />

racesof of maizeinin Latin America.<br />

Through detailed analysis of of these data,<br />

<strong>CIMMYT</strong> scientists have localized several<br />

geographical centers of of origin of of the present<br />

day racesof of maize.<br />

Eachof of these centers has<br />

been located on on the basis of of aa specific combinationof<br />

of basic knobbed positions and sizes<br />

called "chromosome knob complexes".<br />

These complexes include: (1) (1) Mesa Central<br />

Complex, characterizedby by the absenceof<br />

of<br />

knobsininall all the knob forming positions, center<br />

of of originof of this complexisisinin the highlandsof<br />

of<br />

Mexico. (2) Eastern Mexico Complex, characterized<br />

by by the presence of of medium-sized<br />

knobs in in manyof of the knob forming positions.<br />

The centerof of origin of of this complex is is in in the<br />

eastern coastof of Mexico. (3) Zapalote Complex,<br />

having a very special knob combination of of<br />

several sizes in in several of of the knob forming<br />

positions. This complex originated in in southwestern<br />

Mexico, in in the states of of Oaxaca and<br />

Chiapas.<br />

(4) Small Knob Complex, characterized<br />

by the presence of of small knobs in in many<br />

of of the knob forming positions.<br />

Originisisinin the<br />

highlands of of Guatemala. (5) (5) Southern Guatemala<br />

Complex, characterized by by several large<br />

and some small knobs. Center of of origin is is<br />

probably in in the lowlands of of southern Guatemala.<br />

(6) (6) Venezuelan Complex, possesses<br />

large knobsinin several knob forming positions,<br />

primarilyon on the long armsof of the chromosomes.<br />

Geographical center of of origin is is in in some<br />

regionof<br />

of Venezuela.<br />

(7) Andean Complex, characterized<br />

by by the presence of of only two small<br />

knobs; oneon on the long armof of chromosome 7<br />

and the other on on the distal position of of the<br />

long arm of of chromosome 6. 6.<br />

Center of of origin<br />

of of this complexisis locatedinin the highlandsof<br />

of<br />

South America.<br />

Studyof of the Chromosome Knob<br />

Constitutionof of Teosinte<br />

To study the chromosome constitution of of<br />

maize and its its relatives, 14 14 new collections<br />

of of Teosinte were obtained from the States of of<br />

Guanajuato, Mexico, Guerrero and Michoacan<br />

in in Mexico, making aa total of of 39 39 Teosinte collections<br />

among <strong>CIMMYT</strong>'s seed stocks.<br />

In In an an investigation of of the frequency with<br />

which different maize and teosinte races hybridizeinin<br />

nature, several maize races and the<br />

teosintes Chalco (Td and Guerrero (T2)<br />

2 )<br />

were<br />

planted at at the Tepalcingo Experiment Station<br />

in in Mexico. At At flowering time, five pollinations<br />

were made on on shoots of of each maize variety,<br />

using aa mixtureof of equal volumesof of pollenof<br />

of<br />

the variety and Tlo T i • Mixing equal volumes of of<br />

maize and teosinte pollen simulated aa hypothetical<br />

situation in in which such populations<br />

would grow together and coincide in in their<br />

flowering periods. The same procedure was<br />

used in in the case of of T 2 2 to to pollinate the maize<br />

races.<br />

No attempt was made to to use teosinte<br />

populations as as females.<br />

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