V. ashei, 13 % V. constablei and 2 % V. angustifolium. Breeders are finding that the use of V.darrowii has dramatic impacts on fruit quality, and it only takes two or three generations to restorewinter hardiness (Hancock et al, 1995). The complex genetic background of modern breedingpopulations also makes testing of superior genotypes across broad climatic zones imperative tofinding their optimal adaptive zone, particularly for selections of southern and intermediatehighbush families. For example, a few years ago we split our breeding families between Oregonand Michigan, and evaluated them independently at each location (Finn et al., 2003). We used adiverse array of families with varying amounts of southern species blood in their heritage. The elitefamilies that emerged in Michigan where also elite in Oregon, but there were a number of elitefamilies in Oregon that proved poorly adapted to the heat and cold in Michigan. Had we relied onMichigan screens, we would have disregarded some important families.One other important change in highbush breeding is the move towards patenting and licensingblueberry varieties. Today, only the USDA breeding program does not license their varieties, andthey are headed in this direction. This move has come primarily as a means to support furtherbreeding work, as State resources dwindle. Licensing may save some public programs fromextinction due to diminishing state support, but it will also restrict the availability of new varieties.References1. Darnell, R.L., Williamson J.G. (1997) Feasibility of blueberry production in warm climates. Acta Hort.,446, pp. 251-256.2. Finn C.E., Hancock J.F., Mackey T., Serce S. (2003) Genotype x environment interactions in highbushblueberry (Vaccinium sp. L.) families grown in Michigan and Oregon. J. Amer. Soc. Hort. Sci., 128, pp.196-200.3. Hancock J.F. (2006a) Northern highbush breeding. Acta Hort., 715, pp. 37-40.4. Hancock J.F. (2006b) Highbush blueberry breeders. HortScience, 41, pp. 20-21.5. Hancock J.F., Galletta GJ (1995) Dedication: Arlen D. Draper: Blueberry Wizard. Plant BreedingReviews, 13, pp. 1-10.6. Lyrene P.M. (1998) Ralph Sharpe and the Florida blueberry breeding program. In: Cline, WO,Ballington J.R. (eds.). Proceedings of the 8 th North American Blueberry Research and ExtensionWorkers Conf. North Carolina State University, Raleigh. pp. 1-7.7. Lyrene P.M. (2007) Breeding southern highbush blueberries. Plant Breeding Reviews (In press).8. Mainland C.M. (1998) Frederick Coville’s pioneering contributions to blueberry culture and breeding.Proc N Amer Blueberry Workers Conf, Wilmington, NC.9. Sharp R.H., Darrow G.M. (1959) Breeding blueberries for the Florida climate. Proc. Fla. State Hort.Soc., 72, pp. 308-311.FRUIT DEVELOPMENT IN VACCINIUM SPECIESVACCINIUM SUGU AUGěU ATTĪSTĪBALaura JaakolaDepartment of Biology, University of Oulu, POB 3000, FIN-90014 Oulu, Finland,e-mail: laura.jaakola@oulu.fiAbstractFruit development and ripening represent one of the most complex developmental processes inplants. Functionally, the role of fruits is to cover the developing seeds and promote the dispersal ofmature seeds through the production of attractive colour, flavour and aroma compounds. Afterfertilisation, the first phases of fruit development include the division and the expansion of thecells. The ripening phase is initiated after the completed seed maturation. Tissue softening andaccumulation of flavour compounds, aromatic volatiles and pigments occurs during the ripeningphase. The quality of fruits is determined by the different developmental steps via the signallingcascade that is responsible for the metabolic and structural changes during the ripeningphenomenon. Genus Vaccinium is widespread over the world and it includes many economicallyimportant cultivated and wild berry species. Fruits of the Vaccinium species are non-climacteric38
and anatomically they belong to false berries, many-seeded fleshy fruits in which the inferior ovaryalong with floral tube ripens into an edible pericarp. Vaccinium berries are especially rich withphenolic compounds that are known to possess antioxidative properties. This review focuses oncharacteristics of fruit development in both cultivated and the wild Vaccinium species.KopsavilkumsAugĜa attīstība un nogatavošanās ir viens no sarežăītākajiem attīstības procesiem augos.Funkcionāli augĜa uzdevums ir pasargāt jaunattīstītās sēklas un veicināt to nogatavošanos, reizēveicinot arī to iekrāsošanos, atbilstošai garšai un aromātam. Pēc apaugĜošanās, pirmajās augĜaattīstības fāzēs, tiek veicināta šūnu dalīšanās un augšana. AugĜa nogatavošanās fāze sākas pēc tam,kad beidzas sēklu nobriešana. AugĜa audi paliek mīkstāki un garšas un aromātu veidojošosavienojumu un pigmentu uzkrāšanās notiek nogatavošanās fāzes laikā. AugĜa kvalitāti nosakadažādi attīstībā izmantotie signāli, kas atbild par vielmaiĦas un strukturālām pārmaiĦāmnogatavošanās laikā. Vaccinium ăints ir plaši izplatīta visā pasaulē un tajā iekĜautas daudzasekonomiski svarīgas kultivētas un savvaĜas sugas. To augĜi anatomiski pieder pie neīstām ogām –daudzsēklu mīkstajiem augĜiem. Ogas ir īpaši bagātas ar fenola savienojumiem, kam, kā zināms,piemīt antioksidantu īpašības. Šajā rakstā dots augĜa attīstības raksturojums gan kultivētām, gansavvaĜas Vaccinium sugām.Key words: berries, fruit development, Vaccinium.IntroductionFruit development and ripening, typically preceded by successful flower pollination, represent oneof the most complex and important developmental processes in plants. Functionally, the role offruits is to cover the developing seeds and promote the dispersal of mature seeds by frugivoreanimals through the production of attractive colour, flavour and aroma compounds in addition tonutritional value. After fertilisation, the first phases of fruit development include the division andthe expansion of the cells. The ripening phase is initiated after seed maturation has been completed.Tissue softening and accumulation of flavour compounds, aromatic volatiles and pigments occursduring the ripening phase (Brady 1987, Giovannoni 2001, 2004).Fruits can be classified into climacteric and non-climacteric fruits according to the differences inthe respiration rate and the production of the plant hormone ethylene during the ripening phase.According to the structure of the pericarp, fruits are classified as non-dehiscent (fleshy) ordehiscent (dry) fruits. Most fruits develop from a gynoecium that contains one or more carpels. Inpseudocarpic fruits, organs other than the gynoecium (eg. receptacle bracts, floral tube, or theenlarged axis of the inflorescence) participate in the formation of the fruit (Gillaspy et al. 1993,Giovannoni 2004). The genus Vaccinium is widespread over the world with about 450 species ofevergreen and deciduous woody plants varying from dwarf shrubs to trees. Most Vaccinium speciesoriginate from the cooler areas of the Northern hemisphere, although tropical species also exists.Well-known members of the genus are cultivated northern or southern highbush blueberries (V.corymbosum hybrids), lowbush (V. angustifolium) or rabbiteye blueberries (V. ashei Reade) andcranberries (V. macrocarpon), in addition to commercially utilized wild bilberries (V. myrtillus)and lingonberries (V. vitis-idaea). Other better known wild species are European cranberries (V.oxycoccos, V. microcarpum), odon (V. uliginosum) and numerous other especially American wildVaccinium species. The Vaccinium species require soil with low pH and they grow mostabundantly in heaths, bogs and acidic woodlands.Pollination. Most northern highbush blueberries are self-pollinated, meaning that the pollen of thesame individual plant can lead in successful fruit development. However, for many cultivars crosspollination produces higher fruit set and larger fruit. Southern highbush blueberries are onlypartially self-pollinated (Krebs and Hancock 1990). Hokanson and Hancock (2000) tested selffertilityin controlled hand self- and cross-pollinations with individual V. corymbosum, V.angustifolium and V. myrtilloides plants and detected that all three species showed a significantreduction in self fruit set and in the proportion of fertilized ovules that developed into mature seedin self compared to outcross fruit.39
- Page 3 and 4: Conference Organizing CommitteeChai
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each type, and for comparison sampl
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the mean. Kisgyır 1 sample has the
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13. Porpáczy A. (1999) A húsos so
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was medium (0.014 - 0.017 g kg -1 s
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‘Salaspils Ražīgā’. Vigorous
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KopsavilkumsEiropas melleĦu (Vacci
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Figure 2. Chemometric PCA of 32 blu
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References1. Baloga D.W., Vorsa N.,
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obtained from fruits of black choke
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In our opinion, the best estimate a
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cuttings also varies markedly with
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shoots shorter than 10 mm were not
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14. Ostrolucka M.G., Gajdosova A, L
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„Metos RG-350” (http://www.meto
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500480Phenols,mg 100g -146044042040
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SHORT INFORMATION ABOUT THE HISTORY
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Evaluation of cultivars. After the
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the number of pistils (female clone
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Table 2. Number of flowers per harv
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ResultsFirst time upright dieback i
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grew rapidly on PDA at 20 - 24 o C.
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Figure 9. Conidia of Physalospora v
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References1. CABI, EPPO, (1997) Dia
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Results und DiscussionBerries were
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In literature Caruso eds. and Гop
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the total area under a cranberry ma
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Skilled works on development of the
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Tika atrastas dažas būtiskas ats
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appears to maintain a quite low lev
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8. Garkava - Gustavson L.,Persson H