A CONSPECTUS OF THE LICHEN GENUS STEREOCAULON ...

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Journ. Hattori Bot. Lab. No. 43 didymic acid. Distrib. : Costa Rica and Ecuador. Remarks: this is the only species of Stereocaulon known to contain the dibenzofuran didymic acid, and appears to be most closely related to the Hawaiian species St. rubiginosum PERS. (incl. St. roccelloides (TH. FR.) LAMB), having the same peculiar cephalodial structure as the latter. The cephalodial cortex in both these species has in effect the structure of pseudopodetial cortex, being sordid brownish- or isabelline-nubilated, mostly semi-opaque, not gelatinized-conglutinated or palisadic but composed of randomly interwoven or vaguely vertically subparallel, very thick-walled hyphae compacted together but distinct and separate, those on the surface emerging separate and discrete. Algae Scytonema or Stigonema. In the type specimen the apices of some of the sterile pseudopodetia bear terminal, whitish, granulose, pseudosorediate efflorescences, similar to those seen in St. microcarpum MULL. ARG. Morphologically the new species resembles a small state of St. ramulosum (Sw.) RAusc~. or its variety macrocarpurn (RICH.) BAB. Known from 2 collections only; that from Ecuador (FH, MSC) was collected by G. W. PRESCOTT in Prov. Carchi, 1953 (no. An L-5). 102. Stereocaulon dusenii LAMB (n. sp.) Diagn.: Pseudopodetia robusta, rigida, erecta, caespitosa, substrato arcte adnata, 4-7 (-8) cm longa, 1.0-2.5 (4) mm crassa, teretia aut partim leviter complanata, parce modiceve irregulariter ramosa, albida aut cremea, in partibus superioribus corticata, pallio thallino continuo, verrucoso aut rugoso vestita, tantum partibus infimis denudata et ecorticata. Soredia nulla. Phyllocladia cylindrico-coralloidea, valida, crassit. 0.2-0.4 mm, teretia, vulgo nonnihil toruloso-verrucosa, majora ramosa et longit. 4(-5) mm attingentia; pseudopodetia aequaliter vestientia aut passim superne parum evoluta. Cephalodia lateralia, vulgo breviter pedicellata, subglobosa, plumbeo-cinerascentia, olivaceo-cinerea aut subalbida, 1-2 (-2.5) mm lata, scrobiculato-corrugata (ut in St. ramuloso), sacculata, intus laxe contexta, cortice hyalino aut isabellino-nubilato, conglutinato-palisadico. Apothecia terminalia, 1.5-3.0 (-5) mm lata, mox pulvinato-convexa et immarginata, disco fusconigricanti, aetate nonnumquam divisa et discos secundarios proferentia, receptaculo pseudothallino pallido rugosoque reflex0 inferne bene conspicuo. Conus centralis cinerascens et subopacus, laxe aut compacte contextus. Hypothecium parte basali sordide isabellino-nubilatum, parte superiori (50-100 p) incolor et hyalinum. Hymenium 90-100 p altum. Sporae Clnae, rectae, cylindrico-fusiformes, 3-6 (-8)-septatae, longit. 35-60 p, crassit. 4.0-5.5 (-6) p. Pycnidia infra apothecia sita, maculis nigricantibus indicata; conidia filiformia, arcuata, longit. 9-15 p, crassit. 0.3-0.5 p. Atranorinum, acidum fumarprotocetraricum, acidum protocetraricum, acidum perlatolicum et acidum anziaicum continet (K+subfuscescens, PDSminiato-rubescens, saltem in strato medullari). -Simile St. argo HOOK. F. et TAYL. et St. stenospermo (LAMB) LAMB, ab iis materiis chimicis aliisque notis discrepans. Typus: Chile, Magallanes, Isla Desolacibn, Puerto Angosto, altit. ca. 150 m s. m., on rocks, leg. P. DUSBN, 1896 (no. 186) (s, holotypus; CAN, UPS, isotypes). Mat. chim.: atranorin, fumarprotocetraric acid, protocetraric acid, perlatolic acid, anziaic acid. Distrib. : Chile (Magallanes, AysBn, Juan Fernandez) and Argentina (Tierra del Fuego). Remarks: closely related and similar morphologically to St. argus HOOK. F. et TAYL. em. TH. FR., and especially to St. stenospermum (LAMB) LAMB, both of which differ in chemistry and geographical distribution. NYLANDER'S record of St. argus from Chile (1855) probably
I. MACKENZIE LAMB: A Conspectus of Stereocaulon 293 refers to this species. There is a specimen of St. dusenii in s said to be from North America, labelled "Menzies misit. L. rarnulosus Sw. N. America Herb. Swartzii. Stereocaulon ramulosum Achar.", and annotated by TH. FRIES "St. macrocarpum A. Rich.". St. dusenii is sometimes "macrocarpoid" in habitus, with the upper parts of the pseudopodetia more or less destitute of phyllocladia. The cephalodia contain Stigonema or Scytonema. 103. Stereocaulon fronduliferum LAMB (n. sp.) ex MARTIN, Trans. Roy. Soc. New Zealand, Bot. 3 (8): 152 (1966) et MARTIN et CHILD, Lichens of New Zealand : 158 (1972) (nomen nudum). Diagn. : Pseudopodetia arcte affixa, plus minusve erecta, ad 3.5 cm alta (sed saepe minora), ramosa, teretia aut leviter complanata, habitu dorsiventrali (uno latere corticata, glaucescentia, saepe transversim rugosa aut fissa, altero latere decorticata, glabra, ochraceo-albida), ramificatione bilaterali, distincte pinnatifida. Rarnuli phyllocladoidei complanati, simplices aut flabellatim divisi, superficie dorsali glauco-corticati, ventrali pallidiores; longit. 1-2 mm, sed basin pseudopodetiorum versus longiores. Soredia nulla. Cephalodia lateralia, sessilia aut breviter pedicellata, subglobosa, 0.7-1.3 mm lata, primo laevia, dein leviter aut sat distincte foveolata; fuscescentia, subnitida, intus solida (protosacculata), algas Nostocaceas continentia, cortice 40-50 p crasso, incolori aut pallide isabellino, palisadico. Apothecia terminalia, fere ab initio hemisphaerico-convexa et immarginata, tandem fere globosa, 1.0-1.5 (-2.0) mm lata, fusconigricantia, subnitida opacave. Conus centralis laxe contextus, hyphis hyalinis saepe materia viridiflava et granulosa (KHO dissolventi et solutionem aurantiaco-rubescentem effundenti) inspersis. Hypotheciurn pigrnentiferum (fuscum, KHO fere immutatum), ad 150 p crassum. Hymenium 90-110 (-120) p altum. Sporae (6) 8nae, rectae vel subrectae, cylindrico-fusiformes, (4) 5-9 (-10)-septatae, ad septa passim leviter constrictae, longit. 40- 70 (-80) p, crassit. 3.5-4.5 (-5.5) p. Atranorinum, acidum fumarprotocetraricum et fortasse parce acidum protocetraricum continens (K - vel leviter fuscoflavescens, PD intense miniatorubescens). - Species valde peculiaris, phyllocladiis complanatis et flabellato-divisis fere St. foliolosum NYL. simulans. A cel. BABINGTON jam anno 1855 recognitum et nomine St. ramulosum var. compressum appellatum fuit. Typus: New Zealand, exact locality, collector and date not indicated, commun. LOCKE TRAVERS to the Exposition Universelle, Paris, 1889 (PC-HUE, holotypus, named by HUE St. ramulosum var. compressum). Facultative synonyms: Stereocaulon ramulosum var. compressum BAB. in HOOK. F. The Botany of the Antarctic Voyage of H. M. Discovery Ships Erebus and Terror in the Years 1839-1843 . . . Part 11. Flora Novae-Zelandiae. Part ii. Flowerless Plants: 294 (1855). - Stereocaulon ramulosum [subsp. macrocarpum] f. compressum (BAB.) HUE, NOUV. Arch. Mus. Hist. Nat. ser. 3, 2: 245 (1890), nom. illegit., non St. ramulosum f. compressum NYL. ex anno 1860.72 Mat. chim. : atranorin, fumarprotocetraric acid, and probably a trace of protocetraric acid. Distrib.: New Zealand. Unverified records: Colombia (HUE, 1898-1901; LINDAU, 1912), perhaps referring to St. ramulosum f. subcompressum LAMB. Remarks: ZAHLBRUCKNER, Catalogus Lichenum Universalis, vol. I (1922) p. 692, published the combination Stereocaulon compressum, referring to a species of Sphaerophorus, by a lapsus, wrongly ascribing it to LINDSAY. For this reason it seems advisable to use a different name for the present species. Short descriptions (as St. ramulosum var. compressum) were given by TH. FRIES (1857, p. 12, footnote; 1858, p. 26, footnote), NYLANDER (1860, p. 72 A facultative synonym of St. strictum var. compressum (NYL.) LAMB.
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-- f. confluens (MAGN.) FREY 216 --
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-fragile (L.) HOFFM. 327 - fronduli
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-- var. sorediiferum NYL. 226, 312
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- turgescens Nn. 252 - tyroliense (
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A CONSPECTUS OF THE LICHEN GENUS STEREOCAULON ...

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