A CONSPECTUS OF THE LICHEN GENUS STEREOCAULON ...

I. MACKENZIE LAMB: A Conspectus of Stereocaulon 265
Exsicc. ASAH. Lich. Japon. Exs. Fasc. VI (1959) no. 296 (UPS, us). - YOSHIM. Lich. Japon. Exs.
ser. 2 (1972) no. 48 (NICH).
Icon. : SAT6 1941, fig. 23. - ASAH. 1970, fig. 3. - YOSHIM. 1974, fig. 76e; PI. 32, fig. 3 13. - MAGN.
et ZAHLBR. 1945, PI. VII, fig. 1, as Sf. sandwicense.
Mat. chim.: atranorin and lobaric acid.
Distrib. : Japan (Honshu, Hokkaido, Shikoku, Kiushiu); Hawaii. ASAHINA, 1970, states that many
specimens of Sf. octornerellurn have been collected in Honshu and Hokkaido, but only a few in Shikoku
and Kiushiu.
Remarks: this tiny species is saxicolous (very rarely terricolous), firmly attached to rock,
and has a distinct persistent primary thallus consisting of minute, scattered or crowded verruculae
which are not of coralloid form except when giving rise to pseudopodetial initials.
The phyllocladia on the pseudopodetia vary from papillose-verruculose to cylindric-coralloid.
It seems to be always fertile. St. octomerelloides ASAH. was distinguished on the basis of
verruculose phyllocladia on the pseudopodetia, "more delicate appearance and less separated
spores", but these characteristics show all intergradations to the typical condition. St.
sandwicense, formerly considered by us as a distinct variety (LAMB, 1951, LAMB ex FREY 1967),
also intergrades in all characters formerly considered to separate it, and can no longer be
regarded as distinct. St. octomerellum is therefore a somewhat variable species in its morphology
and anatomy. The algae of the cephalodia are usually Stigonema, but in the lectotype
specimen of St. sandwicense from Hawaii (s) they are Calothrix parietina (NAEG.) THUR.
(identified by Dr. F. DROUET), a Cyanophyceous genus not previously reported in Stereocaulon
(cfr. AHMADJIAN, 1967, p. 149/150).
73. Stereocaulon octomerum MULL. ARC.
Flora, 74: 109 (1891); DODGE, Ann. Cryptog. Exot. 2 (2): 115 (1929) (as syn. of St. curtaturn NYL.);
ASAH. J. Jap. Bot. 36 (7): 230 (1961); YOSHIM. Lichen Flora of Japan in Colour : 170 (1974).
Typus: Japan, Honshu, summit of Mt. Ontake, leg. M. MlYos~r, 1890 (no. 113) (G-MULL, holotypus).
Icon.: ASAH. 1961b, fig. 6; 1970, fig. 1A. - YOSHIM. 1974, fig. 76b; P1. 32, fig. 309.
Mat. chim. : atranorin and lobaric acid.
Distrib. : Japan (Honshu, Shikoku, Hokkaido), Formosa, U.S.A. (Alaska).
Remarks: has been confused with St. curtatum NYL., but differs from the latter not only
chemically but also in its taller growth, soon evanescent primary thallus, narrower spores etc.
HUE'S description of "Stereocaulon octomerum", 1898-1901 (1898), p. 249 (sep. 37) obviously
refers to St. curtatum. Although the spores of St. octomerum are normally 7-septate, there
is a good deal of variation; many 3-, 4- or 5-septate spores also occur, and occasionally a
specimen is found with only 34-septate spores. The single Alaskan specimen seen was from
the Kenai Peninsula, leg. H. KROG, 1957 (0); see KROG, 1968, p. 88.
74. Stereocaulon pendulum ASAH.
ex SAT^, J. Jap. Bot. 17: 247 (1941); YOSHIM. Lichen Flora of Japan in Colour: 174 (1974).
Typus: Japan, Honshu, Prov. Kozuke, Mt. Akagi, "ad saxa non calcarea", leg. K. TSUNODA,
1923 (TI, holotypus, not seen by us). Paratypes from Prov. Musashi and Prov. Shinano, sent by Dr.
ASAHINA, were however examined.
48 MAGNUSSON in MAGN. et ZAHLBR. (1944) stated that the exsiccat ZAHLBR. Lich. Rarior. Exs. no.
58, as St. condensaturn, should be referable to St. sandwicense MAGN. But according to our examination
of the examples in PC and UPS, it is St. condensaturn HOFFM. as originally named.