A CONSPECTUS OF THE LICHEN GENUS STEREOCAULON ...

300 Journ. Hattori Bot. Lab. No. 43 1977
branched stipes (undeveloped pseudopodetia) densely covered at their apices with a more or
less continuous crust of dactylaeform-coralloid phyllocladia of minute size (ca. 0.5 mm long,
not over 0.1 mm diam.). Pseudopodetia simple or very sparingly branched, 1.5-2.0 cm high,
0.61.0 mm thick, erect, rigid, decorticate, glabrous, pallid ochraceous. Phyllocladia densely
clothing pseudopodetia from base to apex, cinerascent-whitish, terete-coralloid, simple or
rarely furcate, minute, similar to those on the primary thallus. Cephalodia mostly on primary
thallus, a few also lateral on pseudopodetia; inconspicuous, concolorous with the phyllocladia
or with a slight aeruginose tinge, irregularly subglobose to pulvinate-verrucose, 0.3-1.0 mm
diam., the larger ones becoming divided into more or less discrete verruculae; often with a
few dark spots on the surface due to the Cyanophyceous algae (Stigonema) showing through;
sacculate, loose internally, with cortex 25-60 p thick, colorless and hyaline, highly gelatinized,
with fistulose, branched and anastomosing lumina intricated in various directions (not pali-
sadic). Apothecia terminal, convex-pileate, large, up to 4.5mm diam., dark brown to
blackish-brown, immarginate. Central cone colorless, compact. Hypotheciurn faintly
sordid yellowish in section (not truly pigmented), 45-70 p deep. Hymenium 65-78 p high,
with asci 50-60 p long. Spores seen 4-6 in ascus, cylindric-fusiform, straight, 3-septate,
30-40 x 3.54.0 p. The cephalodia resemble those of St. corticatulum NYL., to which the
species may be related, but it requires further investigation.
1 1 2. Stereocaulon pseudomassartianum LAMB
ex FREY, Bot. Jahrb. Syst. 86: 246 (1967); J. Jap. Bot. 51 (12): 357 (1976).
Typus: Philippines, Luzon, Mountain Province, between Banaue and Mt. Polis, altit. 1060-1212 m
s. m., on soil, leg. A. W. HERRE, 1923 (FH, holotypus; UPS-MAGN, isotypus).
Exsicc.: Lich. Exs. Univ. Colorado Mus. Fasc. VIII (1971) nos. 300, 301, 303 (COLO, FH).
Icon.: FREY 1967, fig. 3: PI. 17, fig. 3. - LAMB 1976, fig. 4 (cephalodia).
Mat. chim. : atranorin, perlatolic acid, sometimes also trace amounts of anziaic acid.
Wrongly stated to contain lobaric acid by LAMB ex FREY (1967).
Distrib. : Philippines, Celebes and New Guinea. Erroneous records: Java and Borneo (LAMB
ex FREY, 1967), =St. halei LAMB.
Remarks: habitually resembles St. massartianum HUE on account of the apothecia borne
on lateral stalks, but immediately distinguishable by the peculiar plate-like or cristate cephalodia,
which are solid-cored and have a dimorphic cortical layer (LAMB, 1976).
1 13. Stereocaulon rubiginosum PERS.
ex GAUDICHAUD in DE FREYCINET, Voyage autour du Monde . . . executB sur les Corvettes de S. M.
1'Uranie et la Physicienne, pendant les annBes 1817, 1818,181 9 et 1820, Botanique: 212 (1826); MAGN.
in MAGN. et ZAHLBR. Ark. Bot. 31A (6): 48 (1944). - Stereocaulon sanguineum DEL. ex GAUDICHAUD,
loc. cit. (illegitimate obligate synonym).
Typus: U.S.A., Hawaii ("Iles Sandwich"), exact locality not stated, on soil, leg. C. GAUDICHAUD
(date ?) (no. 72) (L, holotypus, not seen by us; isotypes studied in PC and H-NYL, also a probable isotype
in UPS-TH FR).
Facultative synonyms: Stereocaulon ramulosum [subsp.]* roccelloides TH. FR. De Stereoc. et
Pilophor. Comment.: 13 (1857); Monogr. Stereoc. et Pilophor.: 330 (sep. 26) (1858); MAGN. in MAGN.
et ZAHLBR. Ark. Bot. 31A (6): 43 (1944). - Stereocaulon roccelloides (TH. FR.) LAMB, J. Jap. Bot. 51
(12): 354, footnote (1976)r4 (Strain 11).
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r4 ZAHLBRUCKNER (192627) and DODGE (1929) cite the taxon as Stereocaulon roccelloides, but this
was a lapsus, not an intentional new combination.