A CONSPECTUS OF THE LICHEN GENUS STEREOCAULON ...

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282 Journ. Hattori Bot. Lab. No. 43 1977 Remarks: characterized by the strongly denuded and ecorticate, ligneous pseudopodetia with smooth, almost polished surface. Appears to be transitional to the var. rnacrocarpum (RICH.) BAB. and the f. elegans TH. FR. Seen from Australia, Tasmania, New Zealand and Chile. Fo. subcompressum LAMB (n. f.) Diagn. : Ramuli phyllocladoidei leviter complanati, flabelliformiter divisi, latere inferiori pallidiores. Ceterum ut in typo. Typus: Tasmania, Mt. Wellington, on rocks, leg. R. A. BASTOW (date ?) (LD, holotypus; FH, isotypus). Remarks: phyllocladia slightly flattened, flabellately branched in one plane and with paler undersides. Otherwise as in the typical species. Rather uncommon; seen from Australia, Tasmania, subantarctic islands of New Zealand, and Chile (incl. Juan Fernandez). Some of the references in the literature to St. ramulosum var. compressum BAB., such as HUE (1898-1901) and LINDAU (1912), perhaps refer to this form of St. ramulosum and not to the true "var. compressum" BAB. (=St. fronduliferum LAMB). Fo. tomentosulum LAMB (n. f.) Diagn. : Pseudopodetiis tomentosulis (tomento tenuissimo, adpresso) dignoscitur; habitu interdum macrocarpoideo (phyllocladiis pseudopodetiorum fertilium superne rarescentibus vel fere nullis). Typus: Colombia, Dept. Santander, W. slope of PBramo Rico, altit. 3600 m s. m., on soil, leg. E. P. KILLIP and A. C. SMITH, 1927 (no. 17733) (s, holotypus; FH, isotypus). Remarks: pseudopodetia more or less continuously clothed with a very thin, adpressedfelted tomentum. Stature and habitus either as in the typical species or more resembling that of var. macrocarpum (RICH.) BAB. with upper parts of fertile pseudopodetia more or less destitute of phyllocladia. Rather common in Colombia and Peru; also seen from Ecuador, Bolivia, Venezuela, Brazil, West Indies (Dominican Republic) and New Zealand. Var. macrocarpum (RICH.) BAB. ex HOOK. F. Botany of the Antarctic Voyage of H. M. Discovery Ships Erebus and Terror in the Years 1839-1843. . .Part 11. Flora Novae-Zelandiae. Part ii. Flowerless Plants: 294 (1855); NYL. Synopsis Methodica Lichenum, 1 (2): 236 (1860). - Stereocaulon macrocarpum RICH. Voyage de Dkouvertes de l'Astrolabe, execut6. . . pendant les annks 18261829 sous le cornmandement de M. Dumont d'u~ille, Botanique, I : 34 (1832); DODGE, Ann. Cryptog. Exot. 2 (2): 139 (l$29)61. - Stereocaulon ramulosum [subsp.]** macrocarpum (RICH.) HUE, NOUV. Arch. Mus. Hist. Nat. ser. 3, 2: 245 (sep. 37) (1890). - Stereocaulon macrocarpum [f.] a. genuinum TH. FR. De Stereoc. et Pilophor. Comment. : 22 (1857). Typus: New Zealand, "loco dicto havre de l'Astrolabe", leg. A. RICHARD (date ?) (PC-HUE, holotypus). This is the specimen cited as type by DODGE, 1929, p. 139, and is labelled: "Echantillon figur6. Nouvelle Zelande. Stereocaulon macrocarpum No. 4. Herbarium Richard"; in red ink has been added, in HUE'S handwriting, the number 234. Facultative synonyms: Stereocaulon proximum [subsp.]** macrocarpoides NYL. Synopsis Methodica Lichenum, 1 (2): 238 (1860). - Stereocaulon proximum var. macrocarpoides (NYL.) NYL Ann. Sci. Nat., Bot. ser. 4, 15: 371 (1861). - Stereocaulon macrocarpoides (NYL.) KREMP. Lichenes, in Reise S. M. The description given by DODGE appears to be a composite one, referring partly to St. implexum TH. FR.
I. MACKENZIE LAMB: A Conspectus of Stereocaulon 283 Fregatte Novara, Bot. Theil, 1 (2): 124 (1870). - Stereocaulon ramulosum var. macrocarpoides (NYL.) MULL. ARG. Bull. Herb. Boissier, 2, Append. I: 22 (1894). Exsicc.: FOLLM. Lich. Exs. Sel. Mus. Hist. Nut. Casselensi Fasc. VII (1975) no. 137 (FH, KASSEL). -Krypt. Exs. Vindob. Cent. XLIV (1963) no. 4339 (FH, LD, s). - V~ZDA, Lich. Sel. Exs. Fasc. XLVIII (1973) no. 1191 (FH). Icon.: RICH. 1832-33, P1. IX, fig. 4, as St. macrocarpum. - DUVIGN. 1956, fig. 10 A, as St. macrocarpum (apothecial structure, asci, spores). Remarks: phyllocladia absent or almost so in upper parts of pseudopodetia, there replaced by a wrinkled and interrupted cortical layer. This character is however somewhat inconstant, and some specimens seem to afford transitions to f. elegans and var. pulvinare. The apothecia are often quite large, up to 4 or 5 mm dim., but sometimes reach this size also in the two other taxa mentioned. Most typically developed in New Zealand and the subantarctic islands, Tierra del Fuego and southern Chile, but material apparently referable here has also been seen from New Caledonia. the New Hebrides, Tahiti, Bolivia, Venezuela and Mexico. Var. pulvinare (DODGE) LAMB (n. comb.) ex D. C. LINDS. Brit. Antarct. Survey Sci. Repts. no. 89: 51 (1974) (comb. inval.). Basionym: Sfereocaulon pulvinare DODGE, Repts. B. A. N. Z. Antarct. Research Exped. 1929- 1931, ser. B, 7: 139 (1948). Typus: New Zealand, South Island, Dunedin, Mac's Valley, on rock, leg. J. S. THOMSON (date?) (no. 9) (DODGE, holotypus). Faculfative synonym (doubtful): Sfereocaulon macquariense DODGE, Nova Hedwigia, 15: 289 (1968) (see Append. 2, p. 323). Remarks: pseudopodetia densely caespitose, crowded and intricated, forming hard and rigid clumps very firmly rooted onto stones or gravelly soil. Perhaps a modification induced by harsh environmental conditions, and possibly not taxonomically distinct from var. macrocarpum. Seen from New Zealand and the subantarctic islands, Australia, Tasmania, southern Chile, Tierra del Fuego, and South Georgia. Fo. crebratum LAMB (n. f.) Diagn.: Habitu maxime compcto et pulvinato, phyllocladiis coralloideis apicalibus congestisque, superficiem crustesam efficientibus. Typus: New Zealand, South Island, Port Chalmers near Dunedin, on rock, leg. J. S. THOMSON (date ?) (no. D. 15, T. 696) (DSIR, holotypus; CAN, MO, isotypes). Remarks: forming pulvinate tufts of extremely compact habitus, with the coralloid phyllocladia apical and very crowded, forming a more or less crustose surface. Usually sterile. Not uncommon in New Zealand and the subantarctic islands; also seen from Tasmania and Juan Fernandez. Var. pilophoroides (TUCK.) LAMB (n. comb.) Basionym: Stereocaulonpilophoroides TUCK. Proc. Amer. Acad. Arts, 6: 265 (1864); DODGE, Ann. Cryptog. Exot. 2 (2): 134 (1929); MAGN. in MAGN. et ZAHLBR. Ark. Bot. 31A (6): 47 (1944); LAMB ex FREY, Bot. Jahrb. Syst. 86: 245 (1967). Typus: U.S.A., Hawaii, without exact locality, leg. W. HILLEBRAND, 1864 (FH-TUCK, holotypus). Facultative synonyms: Stereocaulon denudatum PERS. ex GAUDICHAUD in DE FREYCINET, Voyage autour du Monde . . . executk sur les Corvettes de S. M. 1'Uranie et la Physicienne, pendant les annks 1817, 1818, 1819 et 1820, Botanique: 21 1 (1826) (non St. denudatum FLORKE ex anno 1819). - Stereo-
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-- f. confluens (MAGN.) FREY 216 --
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-fragile (L.) HOFFM. 327 - fronduli
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-- var. sorediiferum NYL. 226, 312
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- turgescens Nn. 252 - tyroliense (
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A CONSPECTUS OF THE LICHEN GENUS STEREOCAULON ...

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