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50<br />

3. MORPHOLOGICAL VARIATION...<br />

plant flowers for the fist time or not. Response to temperature and photoperiod is closely<br />

related to geographic origin of a population and this is probably the reason that the only<br />

L. perenne flowered in the 1 st year of cultivation originated from warm climate (New Zealand).<br />

Date of ear emergence is also a subject of selection according to agronomic management<br />

and would, therefore, reflect the selective pressure during breeding programmes.<br />

Winterhardiness is the second important species determination character. The data<br />

obtained from this study revealed that this character is also quite variable in both species.<br />

It is not surprising either that almost all Scandinavian ecotypes survived the winter<br />

without damage or low winterhardiness in L. perenne from New Zealand. Similarly,<br />

L. multiflorum accessions from Italy rather did not survived the winter while the German<br />

ecotype did quite well. Although significantly greater winterhardiness has been observed in<br />

L. perenne, it seems that this feature strongly depends on geographical origin of a population.<br />

Very high winterhardiness of the Italian ecotype of L. multiflorum (80% of plants recovered<br />

after winter) appears quite surprising. However, Italian ryegrass may behave as<br />

a biennial or even short-lived perennial. In areas with reliable snow cover, it can survive for<br />

five and more years (Cosgrove et al. 1999). Unexpectedly, excellent snow cover in northern<br />

Poland could increase the chances of more southern ecotypes to survive even in comparison<br />

with areas of milder climate but often suffered from little snowing. Moreover the transgressive<br />

segregation observed in many populations suggests that winterhardiness can be increased<br />

above the average level.<br />

On the other hand L. perenne is best adapted to mild-temperate climates. Plants do not<br />

stockpile well and must have good snow cover to survive the winter. This explains why some<br />

L. perenne cultivars were hardly recovered after the winter. It was especially true for Japan<br />

cultivar Kyosato and Polish line Ba012 (about 30% of plants). While the low winterhardiness

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