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3 The New York Years (1931–1953)

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346 11 History and Philosophy of Biology—Mayr’s Third Synthesis<br />

evolutionary synthesis of the 1940s and of post-synthesis developments. In Part<br />

III the work of Mendel’s forerunners, of several species hybridizers and of plant<br />

breeders is explained. <strong>The</strong>se workers did not think in terms of variable populations<br />

describing their results only qualitatively without calculating ratios. <strong>The</strong> species<br />

question and his teacher F. Unger’s interest in variants inspired Gregor Mendel’s<br />

work.Headoptedthemethodofpopulationanalysisandstudiedtensofthousands<br />

of seeds and plants. Although cited several times, his publication of 1866 was<br />

overlooked until it was rediscovered in 1900. Soft and hard inheritance and the<br />

work of A. Weismann and H. de Vries during the late 19th century are discussed in<br />

detail by Mayr as well as the rise of genetics and the chemical basis of inheritance<br />

including the discovery of the structure of DNA in 1953. <strong>The</strong> author exposed<br />

in this volume the historical and philosophical relations between systematics,<br />

evolution and genetics. In addition, general historical issues are discussed in three<br />

introductory chapters and in the epilogue.<br />

Mayr had made particularly good progress with the manuscript during a half<br />

year’s stay in Germany (Würzburg and Tübingen) in 1977. Originally he had<br />

planned to follow up the first volume covering mostly the history of evolutionary<br />

biology (ultimate causations) with a second volume dealing with the history of<br />

functional biology (proximate causations).<br />

“I stayed in Tübingen for about half a year in 1980 working very hard on<br />

the history of physiology and embryology but after about seven months of work<br />

I decided to abandon the project. When there were controversies in 1800 or 1850<br />

in the field of physiology I simply didn’t know enough of physiology to get the<br />

proper feeling for the nature of the controversy nor did I usually know how it<br />

finally came out. I could have produced something strictly descriptive but not the<br />

kind of interpretive story as I had done with the volume devoted to systematics,<br />

evolution, and genetics.”<br />

Despite the fact that he had abandoned the project of a second volume already<br />

in 1980, in the Preface of the published volume of 1982, Mayr still expressed “his<br />

hope to deal with the biology of ‘proximate’ (functional) causations in a later<br />

volume that will cover physiology in all of its aspects, developmental biology, and<br />

neurobiology” (pp. VII–VIII). Evidently the Preface had been written several years<br />

prior to 1982 and was published unchanged.<br />

From the Greeks to Darwin<br />

<strong>The</strong> Greek philosopher Plato (ca. 427–347 B.C.) was a disaster for biology, Mayr<br />

(1982d) concluded. His essentialistic concepts influenced this branch of the natural<br />

sciences adversely for centuries. <strong>The</strong> rise of modern biological thought is, in part,<br />

the emancipation from Platonic thinking. 2 On the other hand, Aristotle (384–<br />

322 B.C.) has made greater contributions to biology than any other thinker before<br />

Darwin. As an empiricist, Aristotle was the founder of the comparative method and<br />

2 <strong>The</strong> history of essentialism is rather complex, because several early thinkers in Europe<br />

had already objected to Platonic idealism a long time before Darwin (Winsor 2006: 3).

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