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Appendices & Glossary - Botanical Research Institute of Texas

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1312 APPENDIX TWENTY-ONE/COMMERCIALLY IMPORTANT TIMBER TREES OF EAST TEXAS<br />

JUGLANS NIGRA L. BLACK WALNUT<br />

(JUGLANDACEAE, WALNUT FAMILY)<br />

Other Common Names: EASTERN BLACK WALNUT, AMERICAN BLACK WALNUT, AMERICAN WALNUT<br />

Form and Size: large tree, 30–40 m tall, rarely to 50 m; trunk straight for about half its height, to 2 m in diameter; the<br />

crown open and rounded at the top.<br />

Leaves: deciduous, alternate, pinnately (=feather-like) compound, composed <strong>of</strong> (9–)15–19(–23) leaflets, the terminal leaflet<br />

small or <strong>of</strong>ten absent; leaflets lanceolate to ovate-lanceolate, (3–)6–15 cm long, 1.5–5.5 cm wide, pointed at the tip,<br />

rounded and uneven at the base, the margins conspicuously toothed, the upper surface yellow-green and glabrous except<br />

for scattered capitate-glandular and other hairs on the midrib, the lower surface paler, with capitate-glandular<br />

hairs, and other hairs and scales; foliage aromatic with distinctive spicy odor, turning yellow in fall.<br />

Buds: terminal buds blunt, broadest at the base and slightly rounded at the top, slightly flattened, pale brown, hairy, 8–10<br />

mm long; lateral buds smaller.<br />

Flowers: unisexual, inconspicuous, male and female flowers on the same tree (plants monoecious), appearing with the<br />

leaves; male flowers in hanging, 5–10 cm long, yellow-green catkins; stamens 17–50 per flower; female flowers usually<br />

1–4 per short spike at the end <strong>of</strong> new growth, each with a single pistil.<br />

Fruits: large, globose, single or in pairs, 3.5–8 cm in diameter, the outer husk thick, green to yellow-green, turning dark<br />

brown at maturity (capable <strong>of</strong> staining skin or clothing), slightly hairy, not splitting open to expose nut, with distinctive<br />

spicy odor; nut globose, 3–4 cm diameter, its shell (visible only when the outer husk <strong>of</strong> fruit is removed) irregularly<br />

and deeply furrowed, the surface between grooves coarsely warty, the kernel 4-lobed, oily, sweet.<br />

Bark: medium to dark gray or brownish, to almost blackish, darkening with age, deeply split into narrow, rough, intersecting<br />

ridges.<br />

Wood: sapwood whitish to yellowish brown (in the trade commonly darkened by streaming or staining to match the<br />

heartwood); heartwood light brown to rich chocolate or purplish brown (the lighter shades from trees grown in the<br />

open); wood with mild characteristic odor; grain straight or irregular (the wavy, curly figures for which this wood is<br />

famous come from burls, crotches, and stump wood); growth rings distinct.<br />

Habitat: stream bottom woods or rich woods on calcium rich soils, with best growth on deep, well-drained, fertile soils.<br />

Range: Pineywoods and northern Gulf Prairies and Marshes west to East Cross Timbers and eastern Edwards Plateau, plus<br />

several locations in northwest <strong>Texas</strong>; widespread in the eastern and central U.S. w to SD and TX.<br />

Principal Uses: this is one <strong>of</strong> the most valuable North American woods and is considered by some to be the finest cabinet<br />

wood; currently, because <strong>of</strong> its rareness, more BLACK WALNUT is being made into veneer, with the veneer used extensively<br />

in furniture and cabinets; the lumber is used for gunstocks (absorbs recoil and does not shrink or warp with age),<br />

furniture, paneling, caskets, and c<strong>of</strong>fins; the ground shells are variously used, including as a non-slip agent for automobile<br />

tires and as a filtering agent for scrubbers in smoke stacks.<br />

Historical Uses: furniture, gunstocks, radio, television, and phonograph cabinets, piano cases, sewing machines,<br />

woodenware, and in pioneer times (when it was abundant) railroad ties and rail fences; Native Americans and settlers<br />

obtained a brown dye from the husk <strong>of</strong> the fruit.<br />

Other Significant Information: although Juglans nigra is not an abundant tree in East <strong>Texas</strong>, it is certainly commercially<br />

important. The wood, which is richly colored, beautifully grained, and easily worked, is so valuable that a single tree<br />

can be worth thousands <strong>of</strong> dollars, and trees are sometimes “rustled.” This species exhibits allelopathy, the inhibition <strong>of</strong><br />

one plant by another via the release <strong>of</strong> chemicals into the environment. The chemical secreted is known as juglone, and<br />

it has been investigated for pharmaceutical and other applications. There are at least 73 patents involving juglone,<br />

demonstrating a range <strong>of</strong> potential applications from a drug useful for AIDS treatment to hair-dyes. Juglone is secreted<br />

primarily by roots but is also found in other tissues (e.g., leaves) and can continue to be secreted for several years after a<br />

tree has been cut. The chemical, technically a naphthoquinone, inhibits the growth <strong>of</strong> many plants, especially vegetables,<br />

though some species are tolerant. Because <strong>of</strong> the possible allelopathic effects, some organic gardeners are careful<br />

concerning the use <strong>of</strong> BLACK WALNUT leaves as mulch. Horses have developed laminitis, a malady <strong>of</strong> the ho<strong>of</strong>, from bedding<br />

in stalls filled with small portions <strong>of</strong> walnut shavings. The nuts have a strong distinctive taste and are used in<br />

baked goods and ice cream. The walnut species now used most frequently commercially is the Old World J. regia L.,<br />

ENGLISH or PERSIAN WALNUT.<br />

Recognition in the Field: alternate, pinnately compound leaves with (9–)15–19(–23) leaflets, the terminal 2–3 leaflets<br />

usually smaller than the middle lateral leaflets; fruits large, round, nut-like, with an indehiscent husk.

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