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Tracing the Source of the Elephant And Hippopotamus Ivory from ...

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differences in <strong>the</strong> frequency <strong>of</strong> species. Ano<strong>the</strong>r culling study by van Hoven (1983) in<br />

Kruger National Park (South Africa) concluded that hippos largely ate grass in proportion<br />

to <strong>the</strong> availability <strong>of</strong> <strong>the</strong> grass, which according to <strong>the</strong> author agrees with an earlier study<br />

in Kruger National Park conducted by Young (1966). Field’s studies, never<strong>the</strong>less, are<br />

more likely accurate, and thus his conclusions more likely correct, since van Hoven only<br />

identified plant species and not <strong>the</strong>ir percentages. However, only differentiating between<br />

browse and graze is necessary for <strong>the</strong> purposes <strong>of</strong> this research, and <strong>the</strong> hippo is<br />

decidedly a grazer, overwhelmingly consuming grasses, some <strong>of</strong> which are likely to be<br />

C4 grasses.<br />

Even so, hippos are known to consume o<strong>the</strong>r types <strong>of</strong> food, particularly in times<br />

<strong>of</strong> environmental stress such as drought or high grazing animal density, which would<br />

limit <strong>the</strong> availability <strong>of</strong> grass. These o<strong>the</strong>r types <strong>of</strong> food include elephant dung (Dudley<br />

1996: 488; Dudley 1998: 58), fruit (i.e. Acacia albida. Eltringham 1999: 79; Dunham<br />

1990), aquatic plants (van Hoven 1983: 48; Field 1970; Mugangu and Hunter 1992;<br />

Eltringham 1999: 79), and even o<strong>the</strong>r animals (Dudley 1996: 486-487, 1998: 58-59;<br />

Eltringham 1999: 82-84).<br />

While it is generally agreed that aquatic plants are not a significant part <strong>of</strong> <strong>the</strong><br />

hippo’s diet, small amounts are consumed. Young (1966) in his Kruger NP study found<br />

traces <strong>of</strong> reeds (Phragmites communis) in <strong>the</strong> stomachs <strong>of</strong> <strong>the</strong> hippos (as reported by van<br />

Hoven 1983: 48). Eltringham (1993, 1999) also reports <strong>from</strong> Field’s research (1970) that<br />

hippos were seen eating a floating plant, <strong>the</strong> Nile cabbage (Pistia stratiotes), and having<br />

witnessed <strong>the</strong> same behavior himself, Eltringham is <strong>of</strong> <strong>the</strong> opinion that <strong>the</strong> hippos<br />

“appeared to be merely toying with <strong>the</strong> plants in a ra<strong>the</strong>r bored, desultory way and it is<br />

97

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