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Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

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<strong>Early</strong><strong>Cretaceous</strong> <strong>Freshwater</strong> <strong>Fish</strong> <strong>Fauna</strong> <strong>in</strong> <strong>Kyushu</strong>, <strong>Japan</strong> 215<br />

Description.<br />

In KMNH VP 100,211, the body is slender and the body depth is conta<strong>in</strong>ed 3.5<br />

times <strong>in</strong> the standard length. The head length is conta<strong>in</strong>ed 3.2 times <strong>in</strong> the standard<br />

length. The snout is round. The outl<strong>in</strong>e of the head dorsal marg<strong>in</strong> is slightly<br />

convex. The outl<strong>in</strong>e from the occipital region to the dorsal orig<strong>in</strong> is almost straight.<br />

The restored outl<strong>in</strong>e of the abdomen is slightly convex (Figs. 78 and 79). The dorsal<br />

f<strong>in</strong> is situated at about the middle of the body. The anal orig<strong>in</strong> is beh<strong>in</strong>d the end of<br />

the dorsal f<strong>in</strong> base. The first dorsal f<strong>in</strong> pterygiophore is <strong>in</strong>serted between the 9th<br />

and 10th abdom<strong>in</strong>al vertebrae. The dorsal f<strong>in</strong> base is short and is conta<strong>in</strong>ed 1.8 times<br />

<strong>in</strong> the anal f<strong>in</strong> base. The pectoral f<strong>in</strong> isshortand does not reach the pelvic <strong>in</strong>sertion.<br />

The caudal f<strong>in</strong> is forked. The number of pr<strong>in</strong>cipal dorsal f<strong>in</strong> rays is 11. There are<br />

12 dorsal f<strong>in</strong> pterygiophores. The number of pr<strong>in</strong>cipal anal f<strong>in</strong> rays is 22. There<br />

are 22 anal f<strong>in</strong> pterygiophores. The pr<strong>in</strong>cipal dorsal and anal f<strong>in</strong> rays are preceded<br />

by one small unbranched accessory ray respectively. Thirteen pectoral f<strong>in</strong> rays are<br />

visible. There are 18 pr<strong>in</strong>cipal caudal f<strong>in</strong> rays (1,8,8,1). The gape of the mouth is<br />

small. The mandible is short and deep. The dentary bears small teeth on the<br />

anterior end of the oral marg<strong>in</strong>. The suture between the dentary and the angloarticular<br />

is visible and the posterior part of the lowerjaw is poorly preserved. The<br />

parasphenoid teeth are absent. The frontal is wide and carries a superficial sensory<br />

canal. The maxilla is wide and slightly curved. The maxilla has a ridge from the<br />

dorsal end to near the ventral end and has wide w<strong>in</strong>gs both sides of the ridge. The<br />

angle between the upper and lower limbs of the preopercle is an obtuse. The<br />

sensory canal runs along the anterior marg<strong>in</strong> of the upper limb. The lower limb of<br />

the preopercle is poorly preserved. Parts of the opercle and preopercle are visible<br />

(Fig. 80). The predorsal scutes are oval <strong>in</strong> shape. The anterior end of each dorsal<br />

scute is round and the posterior end is slightly po<strong>in</strong>ted (Fig. 81). The ventral scutes<br />

are well developed and total 22 <strong>in</strong> number, with 4 scutes below the pectoral f<strong>in</strong>, 11<br />

scutes between the pectoral and pelvic f<strong>in</strong>s, and 7 scutes between the pelvic and anal<br />

f<strong>in</strong>s. Each scute below the pectoral f<strong>in</strong> does not have vertical elements (Fig. 78).<br />

The total number of vertebrae is 36, with 16 caudal vertebrae. The anterior end of<br />

the vertebral column is not visible. The number of abdom<strong>in</strong>al vertebrae was<br />

estimated on the basis ofthe ribs. The number ofribs is 18. Six median supraneu<br />

rals are present. The first supraneural is wide and leaf-shape with a ridge runn<strong>in</strong>g<br />

from the dorsal end to the ventral end. The first preural centrum bears the<br />

cyl<strong>in</strong>drical parhypural with narrow w<strong>in</strong>gs on its basal part. The neural arch and<br />

sp<strong>in</strong>e of the first preural centrum are not visible. There are 6 hypurals. The first<br />

hypural is the largest. The second hypural is about half of the first one <strong>in</strong> width.<br />

The second hypural is fused with the first ural vertebra. There is a space between<br />

the second and the third hypurals. The third to sixth hypurals are slender and<br />

cyl<strong>in</strong>drical (Fig. 82).<br />

In KMNH VP 100,212, the body depth is conta<strong>in</strong>ed 3.7 times <strong>in</strong> the standard

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