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Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

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<strong>Early</strong> <strong>Cretaceous</strong> <strong>Freshwater</strong> <strong>Fish</strong> <strong>Fauna</strong> <strong>in</strong> <strong>Kyushu</strong>, <strong>Japan</strong> 221<br />

Remarks. The characteristic of the caudal skeleton of this new genus, Paraleptolepis<br />

are follows. Anterior two uroneurals extend forward beyond the first ural centrum.<br />

The neural arch and sp<strong>in</strong>e of the first preural centrum are not complete and the<br />

rudimentary neural arch is present on the first preural centrum. Three epurals are<br />

just beh<strong>in</strong>d the neural arch of the first preural centrum.<br />

These characters <strong>in</strong>dicates the relationship with the leptolepid genera, Leptolepis<br />

and Leptolepides, and the elopiform genera, Anaethalion and Pachythrissops. But Paralepto<br />

lepis differs from Anaethalion <strong>in</strong> the characters of the skull. The gape of the mouth is<br />

large and the parasphenoid and maxillary teeth are present <strong>in</strong> Anaethalion (Fig. 83).<br />

Paraleptolepis differs from Pachythrissops <strong>in</strong> the follow<strong>in</strong>g characters. Two rudimentary<br />

neural arches are present on the first ural centrum <strong>in</strong> Pachythrissops (Fig. 84).<br />

Maxillary teeth are present <strong>in</strong> Pachythrissops (Fig. 85). Paraleptolepis is similar to<br />

Leptolepis and Leptolepides <strong>in</strong> the caudal skeleton, but it differs from these genera <strong>in</strong> the<br />

follow<strong>in</strong>g characters. The second uroneural does not reach the preural centrum, one<br />

rudimentary neural arch is present on the first ural centrum, and the maxilla is large<br />

and bears small teeth on its oral marg<strong>in</strong> <strong>in</strong> Leptolepis and Leptolepides. Leptolepis and<br />

Leptolepides were once assigned to the family Leptolepidae of the order Leptolepi<br />

formes, but the group referred to as leptolepids is not considered to be a valid<br />

taxonomic unit (Patterson and Rosen, 1977).<br />

smx 1-2<br />

Fig.83. Anaethalion vidali (Sauvaoe, 1903). Cranium and pectoral girdle <strong>in</strong> left lateral<br />

view. From Forey, 1973.

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