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Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

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244 Yoshitaka Yabumoto<br />

Africa is constituted of about 10 species belong<strong>in</strong>g to 8 families and orders (Patter<br />

son, 1975). The fish fauna of the Ilhas Formation <strong>in</strong> Brazil is composed of 12<br />

species belong<strong>in</strong>g to 7 families and orders (Schaeffer, 1947) and is similar to the<br />

fauna of the Cocobeach Series of Gabon and Equatorial Gu<strong>in</strong>ea. Both faunas are<br />

characterized by the existence of the orders Coelacanthiformes and Aspidorhynchiformes<br />

and an abundance of the Ichthyodectiformes and the Leptolepiformes <strong>in</strong><br />

number of species. The semionotid genus Lepidotes is more abundant and the order<br />

Gonorynchiformes is absent <strong>in</strong> the Ilhas Formation. The orders Amiiformes and<br />

Ichthyodectiformes are common to the Wak<strong>in</strong>o fish fauna and the faunas of the<br />

Cocobeach Series of Gabon and Equatorial Gu<strong>in</strong>ea and the Ilhas Formation.<br />

The orders Coelacanthiformes, Aspidorhynchiformes, and Ellimmichthiiformes<br />

are present <strong>in</strong> the fauna of the Cocobeach Series of Gabon and Equatorial Gu<strong>in</strong>ea<br />

and the Ilhas Formation, but these orders are absent <strong>in</strong> the Wak<strong>in</strong>o fish fauna. The<br />

orders Osteoglossiformes and Clupeiformes which are abundant <strong>in</strong> the Wak<strong>in</strong>o fish<br />

fauna are absent <strong>in</strong> the faunas of the Cocobeach Series of Gabon and Equatorial<br />

Gu<strong>in</strong>ea, and the Ilhas Formation.<br />

The osteoglossiform fish is not recognized <strong>in</strong> the Lower <strong>Cretaceous</strong> fish faunas<br />

except <strong>in</strong> southeastern Ch<strong>in</strong>a and the clupeiform fish is not so abundant <strong>in</strong> the Lower<br />

<strong>Cretaceous</strong> freshwater deposits <strong>in</strong> the world. The Wak<strong>in</strong>o fish fauna is characterized<br />

by an abundance of the orders Osteoglossiformes and Clupeiformes and the absence<br />

of the orders Coccolepiformes, Pholidophoriformes, Macrosemiiformes and Pycnodontiformes,<br />

and the family Huashiidae (Tab. 10).<br />

Conclud<strong>in</strong>g Remarks<br />

1. Comparison of faunal changes of the molluscan fossils and the fish fossils<br />

<strong>in</strong> the Wak<strong>in</strong>o Subgroup.<br />

Y. Ota (1960b) subdivided the Wak<strong>in</strong>o Subgroup distributed <strong>in</strong> the northern<br />

part of <strong>Kyushu</strong> Island <strong>in</strong>to two zones based on molluscan fossils: the lower, Brotiopsis<br />

wak<strong>in</strong>oensis Zone (the lower formation, W,, the First Formation) and the upper,<br />

Viviparus onogoensis-Nakamuranaia (?) sp. cf. N. ch<strong>in</strong>gshanensis Zone. The upper zone is<br />

subdivided <strong>in</strong>to three zones: the middle, upper and uppermost formations which<br />

correspond to the Second (W2), Third (W3) and Fourth (W4) Formations respective<br />

ly on the basis of the distribution of molluscan fossils (Fig.3). In the First<br />

Formation, Brotiopsis wak<strong>in</strong>oensis is most abundant and Plicatounio naktongensis occurs<br />

commonly. "Nippononaia" wak<strong>in</strong>oensis, "N." sengokuensis, Plicatounio naktongensis multiplicatus,<br />

P. triangularis and P. kwanmonensis are very rare <strong>in</strong> the First Formation (Wi).<br />

In the Second Formation (W2), a few molluscan fossils, Viviparus onogoensis and others,<br />

are found from Nogata and Moji <strong>in</strong> northern part of <strong>Kyushu</strong> and Yoshimo <strong>in</strong><br />

Yamaguchi Prefecture. Molluscan fossils of Brotiopsis kobayashii, Viviparus onogoensis<br />

and Sphaerium (?) sp., and Ostracoda are found at Takatsuo <strong>in</strong> Kokura-m<strong>in</strong>ami-ku <strong>in</strong>

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