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Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

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<strong>Early</strong> <strong>Cretaceous</strong> <strong>Freshwater</strong> <strong>Fish</strong> <strong>Fauna</strong> <strong>in</strong> <strong>Kyushu</strong>, <strong>Japan</strong> 243<br />

Prov<strong>in</strong>ce and the Mesoclupea-Huaskia fauna <strong>in</strong> the Zhejiang Prov<strong>in</strong>ce at the early stage<br />

of <strong>Early</strong> <strong>Cretaceous</strong>, and Paralycoplera-Tonghuaichlhys-Huashia fauna <strong>in</strong> the Jil<strong>in</strong>g Prov<br />

<strong>in</strong>ce and the Paralycoptera-P<strong>in</strong>golepis-Huashia fauna <strong>in</strong> the Zhejiang Prov<strong>in</strong>ce at the<br />

early to middle stage of the <strong>Early</strong> <strong>Cretaceous</strong>. The Nipponamia-Aokiichthys fauna is<br />

related to the faunas of the early to middle stage of <strong>Early</strong> <strong>Cretaceous</strong> <strong>in</strong> Ch<strong>in</strong>a on the<br />

basis of its resemblance <strong>in</strong> composition.<br />

Paraleptolepis-Wak<strong>in</strong>oichthys fauna has a family, Chuhsiungichthiidae, <strong>in</strong> common<br />

with the Mesoclupea assemblage, but there is no similarity between the fauna at the<br />

level of genera and species. Diplomystus-Wak<strong>in</strong>oichthys fauna has a genus, Yungkangich<br />

thys, <strong>in</strong> common with the Mesoclupea assemblage, but other members are quite<br />

different. The Paraleptolepis-Wak<strong>in</strong>oichthys and Diplomystus-Wak<strong>in</strong>oichthys faunas differ<br />

from the Mesoclupea assemblage (Tab. 9). The Paraleptolepis-Wak<strong>in</strong>oichthys and Di<br />

plomystus-Wak<strong>in</strong>oichthys faunas are considered to be endemic.<br />

Comparison with other <strong>Cretaceous</strong> freshwater fish faunas <strong>in</strong> the world.<br />

There are several freshwater fish faunas from the Lower <strong>Cretaceous</strong> <strong>in</strong> the world.<br />

In Spa<strong>in</strong>, n<strong>in</strong>eteen species belong<strong>in</strong>g to 13 families and 10 orders are described from<br />

the Lower <strong>Cretaceous</strong> sediments of Las Hoyas and Montsec (Poyato-Ariza and<br />

Wenz, 1990; Sanz et al., 1988). The family Amiidae and the genus Lepidotes are<br />

common to the Las Hoyas and Montesech fauna and the Wak<strong>in</strong>o fish fauna. The<br />

elasmobranch order Hybodontiformes, the primitive halecostome orders Pycnodontiformes<br />

and Macrosemiiformes, the primitive teleostean orders Pholidophoriformes<br />

and Elopiformes, and Coelacanthiformes are present <strong>in</strong> the Las Hoyas and Montsec<br />

fauna, but they are absent <strong>in</strong> the Wak<strong>in</strong>o fish fauna. The orders Ichthyodecti<br />

formes, Osteoglossiformes and Clupeiformes are present <strong>in</strong> the Wak<strong>in</strong>o fish fauna,<br />

but they are absent <strong>in</strong> the Las Hoyas and Montsec fish fauna.<br />

The Wealden freshwater fish fauna <strong>in</strong> Belgium has an order, Amiiformes, and a<br />

genus, Lepidotes, common to the Wak<strong>in</strong>o fish fauna. But the Wealden fish fauna is<br />

characterized by the existence of the chondrostean family Coccolepididae and an<br />

abundance of amiiform fish <strong>in</strong> number of species (Traquair, 1911). The order<br />

Osteoglossiformes which is abundant <strong>in</strong> the Wak<strong>in</strong>o fish fauna is absent <strong>in</strong> the<br />

Wealden fauna.<br />

The fish fauna of Koonwarra <strong>in</strong> Australia is composedof 6 species belong<strong>in</strong>g to 5<br />

families and 5 orders (Waldman, 1971). The order Clupeiformes is common to the<br />

Koonwarra fish fauna and the Wak<strong>in</strong>o fish fauna. However, the chondrostean<br />

family Coccolepididae and the primitive teleostean order Pholidophoriformes are<br />

present <strong>in</strong> the Koonwarra fauna and the existence of the lungfish family Ceratodontidae<br />

is characteristic of the Koonwarra fauna. These families and orders are absent<br />

<strong>in</strong> the Wak<strong>in</strong>o fish fauna. The order Osteoglossiformes is absent <strong>in</strong> the Koonwarra<br />

fauna.<br />

The fish fauna of the Cocobeach Series of Gabon and Equatorial Gu<strong>in</strong>ea <strong>in</strong>

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