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Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

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<strong>Early</strong> <strong>Cretaceous</strong> <strong>Freshwater</strong> <strong>Fish</strong> <strong>Fauna</strong> <strong>in</strong> <strong>Kyushu</strong>, <strong>Japan</strong> 121<br />

Description of the holotype.<br />

The presumed body depth is moderate. The dorsal f<strong>in</strong> is situated at about the<br />

middle of the body. The dorsal f<strong>in</strong> base is longand its length is conta<strong>in</strong>ed 1.4 times<br />

<strong>in</strong> the body depth. There are 14 dorsal f<strong>in</strong> rays with 4 f<strong>in</strong> fulcra. The first f<strong>in</strong><br />

fulcrum of the dorsal f<strong>in</strong> is short and its length is about one third of the second f<strong>in</strong><br />

fulcrum. The second f<strong>in</strong> fulcrum is about half of the third f<strong>in</strong> fulcrum. The third<br />

f<strong>in</strong> fulcrum is about one third of the fourth f<strong>in</strong> fulcrum. The scales of caudal part of<br />

the body are rhombic. The number of scales from the dorsal marg<strong>in</strong> to the ventral<br />

marg<strong>in</strong> at the caudal immediately beh<strong>in</strong>d the dorsal f<strong>in</strong> is 11 to 13.<br />

Remarks. The family Semionotidae of the order Semionotiformes has two genera,<br />

Semionotus and Lepidotes, and the Semionotidae share the presence of dorsal ridge<br />

scales and a large posteriorly directed process on the epiotic (Olsen and McCune,<br />

1991). The presence of dorsal ridge scales and a large posteriorly directed process<br />

on the epiotic are unknown, but the present fossil is similar to Lepidotes yungkangensis<br />

(Chang & Chou, 1974) from <strong>Early</strong> <strong>Cretaceous</strong> deposits of Zhejiang Prov<strong>in</strong>ce <strong>in</strong><br />

Ch<strong>in</strong>a <strong>in</strong> the position of the dorsal and pelvic f<strong>in</strong>s, the number and the form of f<strong>in</strong><br />

fulcra of the dorsal f<strong>in</strong>, the form of scales and the number of scales from the dorsal<br />

marg<strong>in</strong> to the ventral marg<strong>in</strong> at the caudal immediately beh<strong>in</strong>d the dorsal f<strong>in</strong>. But<br />

L. macropterus differs from L. yungkangensis <strong>in</strong> the number of dorsal f<strong>in</strong> rays and the<br />

length of the dorsal f<strong>in</strong> base. The number of dorsal f<strong>in</strong> rays is 14 <strong>in</strong> L. macropterus, 9<br />

<strong>in</strong> L. yungkangensis. The length of the dorsal f<strong>in</strong> base is conta<strong>in</strong>ed 1.4 times <strong>in</strong> the<br />

body depth <strong>in</strong> L. macropterus, 3.0 times <strong>in</strong> L.yungkangensis.<br />

Subdivision Halecomorphi<br />

Order Amiiformes<br />

Family Amiidae<br />

Genus Nipponamia gen. nov.<br />

Diagnosis. It differs from other genera of the Amiidae <strong>in</strong> hav<strong>in</strong>g the comb<strong>in</strong>ation of<br />

follow<strong>in</strong>g characters. The gular plate is almost rhombic <strong>in</strong> the ventral view. The<br />

premaxillary teeth are larger than teeth on the dentary and the number of teeth is 4.<br />

The marg<strong>in</strong> between the condyles of the hyomandibular for the neurocranium and<br />

the opercular process is narrow and deeply concave <strong>in</strong> V-shape. The ventral end of<br />

the hyomandibular is divided <strong>in</strong>to two parts. The number of vertebrae is numerous<br />

and 75 <strong>in</strong>clud<strong>in</strong>g ural centra. The length of vertebral centra is shorter than the<br />

depth. The ural centra are slightly curved upward. Ganoid scales are absent.<br />

Type species.<br />

Nipponamia satoi sp. nov.<br />

Etymology.<br />

The generic name, Nipponamia consists of Nippon mean<strong>in</strong>g <strong>Japan</strong>, and

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