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Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

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<strong>Early</strong><strong>Cretaceous</strong> <strong>Freshwater</strong> <strong>Fish</strong> <strong>Fauna</strong> <strong>in</strong> <strong>Kyushu</strong>, <strong>Japan</strong> 235<br />

meristic characters and the characters of the head and caudal regions, but it differs<br />

from P. kikuchii <strong>in</strong> the proportion. The body is slender and the depth is conta<strong>in</strong>ed<br />

4.7 to 5.5 times <strong>in</strong> the standard length <strong>in</strong> P. elegans, 3.2 to 3.7 times <strong>in</strong> P. kikuchii<br />

(Tab. 6).<br />

Order Gonorynchiformes ?<br />

Family <strong>in</strong>certae sedis<br />

(Figs. 94-95, PI. 59)<br />

Specimen. KMNH VP 100,233, a specimen with its left side exposed. The head<br />

region is miss<strong>in</strong>g. The length from the dorsal orig<strong>in</strong> to the posterior end of the<br />

hypural is 16.2 mm. The estimated standard length is 32.5 mm.<br />

Locality. M<strong>in</strong>amigaoka (KM-1), Kokura-kita-ku (Kokura Northern Ward), Kita<br />

kyushu City, Fukuoka Prefecture, <strong>Japan</strong>.<br />

Horizon. The Third Formation (the upper formation, W3, correlated to the Lower<br />

Wakamiya Formation) of the Wak<strong>in</strong>o Subgroup <strong>in</strong> the Kwanmon Group, the Lower<br />

<strong>Cretaceous</strong>.<br />

Description.<br />

The body is slender. The body depth is conta<strong>in</strong>ed 7.6 times <strong>in</strong> the estimated<br />

standard length. The outl<strong>in</strong>e from the occipital region to the dorsal orig<strong>in</strong> is<br />

straight. The dorsal f<strong>in</strong> is situated at about the middle of the body. The pelvic f<strong>in</strong><br />

is large and is situated below the dorsal f<strong>in</strong> (Fig. 94, PI. 59). The first dorsal f<strong>in</strong><br />

pterygiophore is <strong>in</strong>serted between the neural sp<strong>in</strong>es of the eighth and n<strong>in</strong>th abdomi<br />

nal vertebrae. The dorsal f<strong>in</strong> is large and the base is longer than the anal f<strong>in</strong> base.<br />

The anal f<strong>in</strong> base is conta<strong>in</strong>ed 1.6 times <strong>in</strong> the dorsal f<strong>in</strong> base. The pectoral f<strong>in</strong> is<br />

large and extends below the dorsal orig<strong>in</strong>. The orig<strong>in</strong> of the anal f<strong>in</strong> is beh<strong>in</strong>d the<br />

end of the dorsal f<strong>in</strong> base. The caudal f<strong>in</strong> is forked. The number of pr<strong>in</strong>cipal<br />

dorsal f<strong>in</strong> rays is 12. There are 12 dorsal f<strong>in</strong> pterygiophores. The number of<br />

pr<strong>in</strong>cipal anal f<strong>in</strong> rays is 11. Thereare 11 anal f<strong>in</strong> pterygiophores. Twelve pectoral<br />

f<strong>in</strong> rays are visible. Seven pelvic f<strong>in</strong> rays are visible. There are 19 pr<strong>in</strong>cipal caudal<br />

f<strong>in</strong> rays (1,9,8,1). The total number of vertebrae is 35, with 18 caudal vertebrae.<br />

There is a series of median supraneurals beg<strong>in</strong>n<strong>in</strong>g immediately beh<strong>in</strong>d the head and<br />

end<strong>in</strong>g before the dorsal orig<strong>in</strong>. Each supraneural is cyl<strong>in</strong>drical. The neural arch<br />

and sp<strong>in</strong>e of the first preural centrum are complete. There are 6 hypurals. The<br />

first hypural is the largest. The second hypural is about half of the first one <strong>in</strong> width<br />

and does not extend to the first ural centrum. There is a space between the second<br />

and third hypurals. There are two ural centra. The second ural centrum bears<br />

three hypurals (HYU3-5). One uroneural is visible and extends forward to the first

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