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Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

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40-42<br />

2.9<br />

<strong>Early</strong> <strong>Cretaceous</strong> <strong>Freshwater</strong> <strong>Fish</strong> <strong>Fauna</strong> <strong>in</strong> <strong>Kyushu</strong>, <strong>Japan</strong> 137<br />

Table 2.<br />

Comparison of characters <strong>in</strong> the speciesof the family Chuhsiungichthyiidae nov.<br />

Chuhsiungichthysyanagidai<br />

sp. nov.<br />

C.japonicus sp. nov. C. tsangl<strong>in</strong>gensis Mesoclupea shoivchangensis<br />

SL 168.2 mm 217.0-220.0 imm 50-140 mm 40-170 mm<br />

counts<br />

D. 20-21 21 21-23 14<br />

A. 32 39 39-40 46<br />

P. 14+ 18 15 12<br />

Vertebrae 40 —<br />

ratio<br />

53-54<br />

SL/BD 3.6 2.4-2.5 3.0-3.7 2.7-3.4<br />

SL/HL 4 —<br />

3.8<br />

AB/DB 1.4 1.6 1.9 4.2<br />

A., anal f<strong>in</strong> rays; AB, anal f<strong>in</strong> base; BD, body depth; D., dorsal f<strong>in</strong> rays; DB, dorsl f<strong>in</strong> base; HL,<br />

head length; P|, pectoral f<strong>in</strong> rays; SL, standard length.<br />

Remarks. Although the present fossils are poorly preserved <strong>in</strong> head region, it<br />

clearly shows a character to be a member of the order Ichthyodectiformes by the<br />

uroneurals cover<strong>in</strong>g the lateral surfaces of the preural centra (Fig. 21). The<br />

Ichthyodectiformes conta<strong>in</strong>s 13 or 14 genera from Upper Jurassic to Lower Cre<br />

taceous deposits of all cont<strong>in</strong>ents <strong>in</strong> the world (Patterson and Rosen, 1977).<br />

Among them, the present new species is considered to belong to the genus Chuhsiung<br />

ichthys from Upper <strong>Cretaceous</strong> of Yunnan Prov<strong>in</strong>ce (Lew, 1974) because this species<br />

is closest to Chuhsiungichthys tsangl<strong>in</strong>gensis Lew, 1974 (Fig. 22) <strong>in</strong> the proportion and<br />

meristic characters with the exception of number of anal f<strong>in</strong> rays (D. 21-23; A. 39-<br />

40; V. 40-42 <strong>in</strong> C. tsangl<strong>in</strong>gensis) (Tab. 2). This species differs from Mesoclupea,<br />

which has a s<strong>in</strong>gle species M. shoivchangensis P<strong>in</strong>g & Yen, 1933 (Fig. 23), from Late<br />

Jurassic of Zhejiang Prov<strong>in</strong>ce and Chekiang Prov<strong>in</strong>ce (P<strong>in</strong>g and Yen, 1933; Chang,<br />

1963;Chang and Chou, 1974, 1977; Chang and Chow, 1986) <strong>in</strong> the position and the<br />

size of the dorsal f<strong>in</strong> (the dorsal f<strong>in</strong> is small and the dorsal base is situated above the<br />

middle of anal f<strong>in</strong> base <strong>in</strong> Mesoclupea) and the meristic characters (D. 14; A. 46; V.<br />

53-54 <strong>in</strong> Mesoclupea and D. 20-21; A. 32; V. 40 <strong>in</strong> this species). Chuhsiungichthys<br />

yanagidai differs from Chuhsiungichthysjaponicus with more slender body (the body depth<br />

is conta<strong>in</strong>ed 2.4 to 2.5 times <strong>in</strong> the standard length <strong>in</strong> C.japonicus) and <strong>in</strong> the number<br />

of anal f<strong>in</strong> rays (Tab. 2).<br />

Chuhsiungichthys japonicus sp. nov.<br />

(Figs. 24-25, PI. 41)<br />

Diagnosis. This species differs from other species of the genus Chuhsiungichthys <strong>in</strong><br />

hav<strong>in</strong>g the comb<strong>in</strong>ationof follow<strong>in</strong>g characters. The dorsal f<strong>in</strong> is large. The dorsal

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