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Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

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<strong>Early</strong><strong>Cretaceous</strong> <strong>Freshwater</strong> <strong>Fish</strong> <strong>Fauna</strong> <strong>in</strong> <strong>Kyushu</strong>, <strong>Japan</strong> 173<br />

vertebrae. The number of vertebra is 34 to 35 <strong>in</strong> A. uyenoi, 44 <strong>in</strong> Y. hsitanensis. A.<br />

uyenoi is closest to Aokiichthys otai <strong>in</strong> hav<strong>in</strong>g almost same meristic characters and<br />

almost same proportion with the exception of the relative position of the median f<strong>in</strong>s<br />

(Tab. 3). The orig<strong>in</strong> of the dorsal f<strong>in</strong> is vertical with the fourth anal f<strong>in</strong> pteryg<br />

iophore<strong>in</strong> A. uyenoi. The orig<strong>in</strong>ofdorsal f<strong>in</strong> is vertical with the anal orig<strong>in</strong> <strong>in</strong> A. otai.<br />

Aokiichthys praedorsalis sp. nov.<br />

(Figs. 48-50, pis. 48-49)<br />

Diagnosis. It differs from other species of the genus Aokiichthys <strong>in</strong> hav<strong>in</strong>g the<br />

comb<strong>in</strong>ation of follow<strong>in</strong>g characters. The dorsal f<strong>in</strong> is large. The anal orig<strong>in</strong> is<br />

beh<strong>in</strong>d the dorsal orig<strong>in</strong> and about vertical with the 7th dorsal f<strong>in</strong> pterygiophore<br />

(the middle of the dorsal f<strong>in</strong> base). The length of dorsal f<strong>in</strong> base is conta<strong>in</strong>ed 1.2<br />

times <strong>in</strong> the length of anal f<strong>in</strong> base. The number of dorsal f<strong>in</strong> rays is 11 to 13. The<br />

number of anal f<strong>in</strong> rays is 15 to 17. The parasphenoid teeth are larger than those of<br />

the premaxilla and the dentary. The length of lower jaw is conta<strong>in</strong>ed 1.5 times <strong>in</strong><br />

the head length. The known size is 57.7 to 146.7 mm.<br />

Holotype. KMNH VP 100,155, an almost complete specimen, but a dorsal part of<br />

caudal peduncle is miss<strong>in</strong>g. The head and caudal skeletons are poorly preserved.<br />

The standard length is 146.7 mm.<br />

Paratypes. KMNH VP 100,156, an anterior part of the body. The length from the<br />

anterior end of the premaxilla to the posterior end of the 20th vertebra is 82 mm.<br />

KMNH VP 100,157, an almost complete specimen with its right side exposed. The<br />

standard length is 61.8 mm. KMNH VP 100,158, an almost complete specimen,<br />

with its left side exposed. The standard length is 57.7 mm.<br />

Locality. Tokuriki (TO-1), Kokura-m<strong>in</strong>ami-ku (Kokura Southern Ward), Kitakyu<br />

shu City, Fukuoka Prefecture, <strong>Japan</strong>.<br />

Horizon. The First Formation (the lower formation, Wj, correlated to the Sengoku<br />

Formation) of the Wak<strong>in</strong>o Subgroup <strong>in</strong> the Kwanmon Group, the Lower <strong>Cretaceous</strong>.<br />

Etymology: The species name, praedorsalis, means anterior position of the dorsal f<strong>in</strong>,<br />

which refers to the fact that the dorsal orig<strong>in</strong> is situated before the anal f<strong>in</strong> <strong>in</strong> this<br />

species.<br />

Description of the holotype.<br />

The body is deep. The body depth is conta<strong>in</strong>ed 2.8 times <strong>in</strong> the standard

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