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The Origin and Evolution of Mammals - Moodle

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164 THE ORIGIN AND EVOLUTION OF MAMMALS<br />

(a)<br />

(b)<br />

(c)<br />

Tinodon<br />

Woutersia<br />

Bondesius<br />

pa.d<br />

pr.d<br />

me.d<br />

in North America <strong>and</strong> Europe, is a typical example.<br />

Hu et al. (1997) described a largely complete spalacotheriid<br />

skeleton from the Early Cretaceous Yixian<br />

Formation <strong>of</strong> northeastern China (Zhou et al. 2003).<br />

(d)<br />

(e)<br />

M 3<br />

M 4 M 5<br />

M 3 M 4 M 5 M 6<br />

Zhangheotherium<br />

Figure 5.14 Symmetrodontans. (a) Upper molar <strong>of</strong> Eurylambda in occlusion with two lower molars <strong>of</strong> the closely related Tinodon, in<br />

bucco-lingual <strong>and</strong> occlusal views (Crompton <strong>and</strong> Jenkins 1967). (b) Isolated upper <strong>and</strong> lower molars <strong>of</strong> Woutersia butleri in occlusal <strong>and</strong> lingual<br />

views (Sigogneau-Russell <strong>and</strong> Hahn 1995). (c) Isolated right lower molar <strong>of</strong> Bondesius ferox in occlusal, labial, <strong>and</strong> lingual viens (Bonaparte 1990).<br />

(d) Lower jaw <strong>of</strong> Zhangheotherium quinquecuspidens in medial view, with enlarged views <strong>of</strong> upper <strong>and</strong> lower molars. Length <strong>of</strong> jaw approx. 2.5 cm<br />

(Hue et al. 1997). (e) Skeleton <strong>of</strong> Zhangheotherium quinquecuspidens as preserved. Maximum length approx. 13 cm (Hu et al. 1997).<br />

Named Zhangeotherium (Fig. 5.14(d) <strong>and</strong> (e)), it has<br />

added enormously to the detailed knowledge <strong>of</strong> a<br />

basal holotherian, <strong>and</strong> reveals a number <strong>of</strong> additional<br />

characters shared with the later holotherians.

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