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The Origin and Evolution of Mammals - Moodle

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(Fig. 3.2(f)), is actually an ophiacodontid. It is a small<br />

pelycosaur with a skull about 9 cm long <strong>and</strong> a body<br />

length <strong>of</strong> around 50 cm including the tail, that was<br />

discovered in fossilised lycopod tree stumps <strong>of</strong><br />

Florence, Nova Scotia, in Pennsylvanian deposits<br />

dated at about 315 Ma. Its long, slender snout bears a<br />

row <strong>of</strong> little, sharply pointed teeth, one <strong>of</strong> which is<br />

distinctly caniniform. <strong>The</strong> postcranial skeleton <strong>of</strong><br />

(a)<br />

(d)<br />

Edaphosaurus<br />

(e)<br />

(b)<br />

EVOLUTION OF MAMMAL-LIKE REPTILES 23<br />

Archaeothyris is not yet very well known. <strong>The</strong> generally<br />

similar, but relatively much larger <strong>and</strong> longer<br />

snouted Ophiacodon (Fig. 3.5(a)) is one <strong>of</strong> the best<br />

known <strong>of</strong> all pelycosaurs. It appears in the latest<br />

Upper Carboniferous Stephanian <strong>of</strong> North America<br />

<strong>and</strong> remains common throughout the Early Permian,<br />

when some species achieved a very large size, up to<br />

about 300 cm in length. <strong>The</strong>re are several indications<br />

(c)<br />

Edaphosaurus<br />

Ophiacodon<br />

Glaucosaurus<br />

Ianthasaurus<br />

Figure 3.5 Ophiacodontid <strong>and</strong> edaphosaurid pelycosaurs. (a) reconstructed skeleton <strong>of</strong> Ophiacodon retroversus (Romer <strong>and</strong> Price 1940).<br />

(b) Incomplete skull <strong>of</strong> Glaucosaurus megalops (Modesto 1994). (c) Specimen <strong>of</strong> Ianthasaurus hardestii showing cross-pieces on neural spines, <strong>and</strong><br />

simple dentition (Reisz <strong>and</strong> Berman 1986). (d) Reconstructed skeleton <strong>of</strong> Edaphosaurus boanerges (Romer <strong>and</strong> Price 1940). (e) Skull <strong>of</strong><br />

Edaphosaurus boanerges in dorsal, ventral <strong>and</strong> lateral views, <strong>and</strong> internal view <strong>of</strong> lower jaw (Modesto 1995).

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